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Vestibulospinal Tract
The vestibulospinal tract is a neural tract in the central nervous system. Specifically, it is a component of the extrapyramidal system and is classified as a component of the medial pathway. Like other descending motor pathways, the vestibulospinal fibers of the tract relay information from nuclei to motor neurons. The vestibular nuclei receive information through the vestibulocochlear nerve about changes in the orientation of the head. The nuclei relay motor commands through the vestibulospinal tract. The function of these motor commands is to alter muscle tone, extend, and change the position of the limbs and head with the goal of supporting posture and maintaining balance of the body and head. Classification The vestibulospinal tract is part of the "extrapyramidal system" of the central nervous system. In human anatomy, the extrapyramidal system is a neural network located in the brain that is part of the motor system involved in the coordination of movement. The system i ...
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Neural Tract
A nerve tract is a bundle of nerve fibers (axons) connecting nuclei of the central nervous system. In the peripheral nervous system this is known as a nerve, and has associated connective tissue. The main nerve tracts in the central nervous system are of three types: association fibers, commissural fibers, and projection fibers. A tract may also be referred to as a commissure, decussation, pathway or fasciculus. A commissure connects the two cerebral hemispheres at the same levels, while a decussation connects at different levels (crosses obliquely). Types The nerve fibers in the central nervous system can be categorized into three groups on the basis of their course and connections. Different tracts may also be referred to as ''projections'' or ''radiations''. Association fibers The tracts that connect cortical areas within the same hemisphere are called association tracts. Long association fibers connect different lobes of a hemisphere to each other whereas short assoc ...
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Reticulospinal Tract
The reticular formation is a set of interconnected nuclei that are located throughout the brainstem. It is not anatomically well defined, because it includes neurons located in different parts of the brain. The neurons of the reticular formation make up a complex set of networks in the core of the brainstem that extend from the upper part of the midbrain to the lower part of the medulla oblongata. The reticular formation includes ascending pathways to the cortex in the ascending reticular activating system (ARAS) and descending pathways to the spinal cord via the reticulospinal tracts. Neurons of the reticular formation, particularly those of the ascending reticular activating system, play a crucial role in maintaining behavioral arousal and consciousness. The overall functions of the reticular formation are modulatory and premotor, involving somatic motor control, cardiovascular control, pain modulation, sleep and consciousness, and habituation. The modulatory functions are prim ...
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Lateral Funiculus
The most lateral of the bundles of the anterior nerve roots is generally taken as a dividing line that separates the anterolateral system into two parts. These are the anterior funiculus, between the anterior median fissure and the most lateral of the anterior nerve roots, and the lateral funiculus (or lateral column) between the exit of these roots and the posterolateral sulcus. The lateral funiculus transmits the contralateral corticospinal and spinothalamic tracts. A lateral cutting of the spinal cord results in the transection of both ipsilateral posterior column and lateral funiculus and this produces Brown-Séquard syndrome Brown-Séquard syndrome (also known as Brown-Séquard's hemiplegia, Brown-Séquard's paralysis, hemiparaplegic syndrome, hemiplegia et hemiparaplegia spinalis, or spinal hemiparaplegia) is caused by damage to one half of the spinal cord, i.e. hem ....Kaplan Qbook - USMLE Step 1 - 5th edition - page References Central nervous system {{ ...
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Efferent Nerve Fiber
Efferent nerve fibers refer to axonal projections that ''exit'' a particular region; as opposed to afferent projections that ''arrive'' at the region. These terms have a slightly different meaning in the context of the peripheral nervous system (PNS) and central nervous system (CNS). The efferent fiber is a long process projecting far from the neuron's body that carries nerve impulses away from the central nervous system toward the peripheral effector organs (mainly muscles and glands). A bundle of these fibers is called an efferent nerve (if it connects to muscles, then it is a motor nerve). The opposite direction of neural activity is afferent conduction, which carries impulses by way of the afferent nerve fibers of sensory neurons. In the nervous system there is a "closed loop" system of sensation, decision, and reactions. This process is carried out through the activity of sensory neurons, interneurons, and motor neurons. In the CNS, afferent and efferent projections ...
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Interneurons
Interneurons (also called internuncial neurons, relay neurons, association neurons, connector neurons, intermediate neurons or local circuit neurons) are neurons that connect two brain regions, i.e. not direct motor neurons or sensory neurons. Interneurons are the central nodes of neural circuits, enabling communication between sensory or motor neurons and the central nervous system (CNS). They play vital roles in reflexes, neuronal oscillations, and neurogenesis in the adult mammalian brain. Interneurons can be further broken down into two groups: local interneurons and relay interneurons. Local interneurons have short axons and form circuits with nearby neurons to analyze small pieces of information. Relay interneurons have long axons and connect circuits of neurons in one region of the brain with those in other regions. However, interneurons are generally considered to operate mainly within local brain areas. The interaction between interneurons allow the brain to perform compl ...
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Lateral Vestibulospinal Tract
The lateral vestibulospinal tract is one of the descending spinal tracts of the ventromedial funiculus. The lateral part of the vestibulospinal tract is the major portion and is composed of fibers originating in the lateral, superior, and inferior vestibular nuclei (primarily the lateral). It projects ipsilaterally down to the lumbar region of the spinal cord. There it helps to maintain an upright and balanced posture by stimulating extensor motor neurons in the legs. It also innervates muscles of the trunk, thus additionally aiding in body posture. The lateral vestibular nuclei receive input from cerebellum, particularly the vestibulocerebellum, or the flocculi and nodulus. The cerebellum aids in coordinating postural adjustments. See also *Decerebrate posturing Abnormal posturing is an involuntary flexion or extension of the arms and legs, indicating severe brain injury. It occurs when one set of muscles becomes incapacitated while the opposing set is not, and an ex ...
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Tectospinal Tract
In humans, the tectospinal tract (or colliculospinal tract) is a nerve tract that coordinates head and eye movements. This tract is part of the extrapyramidal system and connects the midbrain tectum, and cervical regions of the spinal cord. It is responsible for motor impulses that arise from one side of the midbrain to muscles on the opposite side of the body (contralateral). The function of the tectospinal tract is to mediate reflex postural movements of the head in response to visual and auditory stimuli. The portion of the midbrain from where this tract originates is the superior colliculus, which receives afferents from the visual nuclei (primarily the oculomotor nuclei complex), then projects to the contralateral (decussating dorsal to the mesencephalic duct) and ipsilateral portion of the first cervical neuromeres of the spinal cord, the oculomotor and trochlear nuclei in the midbrain and the abducens nucleus in the caudal portion of the pons. The tract descends to t ...
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Medial Longitudinal Fasciculus
The medial longitudinal fasciculus (MLF) is an area of crossed over tracts, on each side of the brainstem. These bundles of axons are situated near the midline of the brainstem. They are made up of both ascending and descending fibers that arise from a number of sources and terminate in different areas, including the superior colliculus, the vestibular nuclei, and the cerebellum. It contains the interstitial nucleus of Cajal, responsible for oculomotor control, head posture, and vertical eye movement. The medial longitudinal fasciculus is the main central connection for the oculomotor nerve, trochlear nerve, and abducens nerve. It carries information about the direction that the eyes should move. Lesions of the medial longitudinal fasciculus can cause nystagmus and diplopia, which may be associated with multiple sclerosis, a neoplasm, or a stroke. Structure The medial longitudinal fasciculus is an area of crossed over tracts, on each side of the brainstem. It is medial, ...
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Medial Vestibular Nucleus
The medial vestibular nucleus (Schwalbe nucleus) is one of the vestibular nuclei. It is located in the medulla oblongata. Lateral vestibulo-spinal tract (lateral vestibular nucleus “Deiters”)- via ventrolateral medulla and spinal cord to ventral funiculus (lumbo-sacral segments). ..Ipsilaterally for posture Medial vestibulo-spinal tract (medial, lateral, inferior, vestibular nuclei), bilateral projection via descending medial longitudinal fasciculus to cervical segments. DESCENDING MLF..Bilaterally for head/neck/eye movements It is one of the nuclei that corresponds to CN VIII, corresponding to the vestibular nerve, which joins with the cochlear nerve. It receives its blood supply from the Posterior Inferior Cerebellar Artery, which is compromised in the lateral medullary syndrome. See also * Vestibular nerve * Vestibular system References External links

* https://web.archive.org/web/20111109232230/http://www.neuroanatomy.wisc.edu/virtualbrain/BrainStem/13VNAN.html ...
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Medial Vestibulospinal Tract
The medial vestibulospinal tract is one of the descending spinal tracts of the ventromedial funiculus of the spinal cord. It is found only in the cervical spine and above. The medial part of the vestibulospinal tract is the smaller part, and is primarily made of fibers from the medial vestibular nucleus. It projects bilaterally down the spinal cord and triggers the ventral horn of the cervical spinal circuits, particularly controlling lower motor neurons associated with the spinal accessory nerve (CN XI). Additionally, the pathway projects superiorly to the paramedian pontine reticular formation The paramedian pontine reticular formation, also known as PPRF or paraabducens nucleus, is part of the pontine reticular formation, a brain region without clearly defined borders in the center of the pons. It is involved in the coordination of eye ..., indirectly innervating the nuclei of CN VI and III. Through this superior projection, the medial vestibulospinal tract is involved in " ...
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Upper Motor Neuron
Upper motor neurons (UMNs) is a term introduced by William Gowers in 1886. They are found in the cerebral cortex and brainstem and carry information down to activate interneurons and lower motor neurons, which in turn directly signal muscles to contract or relax. UMNs in the cerebral cortex are the main source of voluntary movement. They are the larger pyramidal cells in the cerebral cortex. There is a type of giant pyramidal cell called ''Betz cells'' and are found just below the surface of the cerebral cortex within layer V of the primary motor cortex. The cell bodies of Betz cell neurons are the largest in the brain, approaching nearly 0.1 mm in diameter. The primary motor cortex, or precentral gyrus, is the most posterior gyrus of the frontal lobe and lies anterior to the central sulcus. The pyramidal cells of the precentral gyrus are also called upper motor neurons. The fibers of the upper motor neurons project out of the precentral gyrus ending in the brainstem ...
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Extraocular Muscles
The extraocular muscles (extrinsic ocular muscles), are the seven extrinsic muscles of the human eye. Six of the extraocular muscles, the four recti muscles, and the superior and inferior oblique muscles, control movement of the eye and the other muscle, the levator palpebrae superioris, controls eyelid elevation. The actions of the six muscles responsible for eye movement depend on the position of the eye at the time of muscle contraction. Structure Since only a small part of the eye called the fovea provides sharp vision, the eye must move to follow a target. Eye movements must be precise and fast. This is seen in scenarios like reading, where the reader must shift gaze constantly. Although under voluntary control, most eye movement is accomplished without conscious effort. Precisely how the integration between voluntary and involuntary control of the eye occurs is a subject of continuing research."eye, human."Encyclopædia Britannica from Encyclopædia Britannica 2006 Ulti ...
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