Tetanurans
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Tetanurans
Tetanurae (/ˌtɛtəˈnjuːriː/ or "stiff tails") is a clade that includes most theropod dinosaurs, including megalosauroids, allosauroids, tyrannosauroids, ornithomimosaurs, compsognathids and maniraptorans (including birds). Tetanurans are defined as all theropods more closely related to modern birds than to '' Ceratosaurus'' and contain the majority of predatory dinosaur diversity. Tetanurae likely diverged from its sister group, Ceratosauria, during the late Triassic. Tetanurae first appeared in the fossil record by the Early Jurassic about 190 mya and by the Middle Jurassic had become globally distributed. The group was named by Jacques Gauthier in 1986 and originally had two main subgroups: Carnosauria and Coelurosauria, the clade containing birds and related dinosaurs such as compsognathids, tyrannosaurids, ornithomimosaurs, and maniraptorans. The original Carnosauria was a polyphyletic group including any large carnivorous theropod. Many of Gauthier's carnosaurs, su ...
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Allosauroidea
Allosauroidea is a superfamily or clade of theropod dinosaurs which contains four family (biology), families — the Metriacanthosauridae, Allosauridae, Carcharodontosauridae, and Neovenatoridae. Allosauroids, alongside the family Megalosauroidea, were among the apex predators that were active during the Middle Jurassic to Late Cretaceous periods. The most famous and best understood allosauroid is the North American genus ''Allosaurus''. The oldest-known allosauroid, ''Shidaisaurus jinae'', appeared in the early Middle Jurassic about 174 million years ago (Earliest Aalenian stage) of China. The last known definitive surviving members of the group died out around 89 million years ago in Asia (''Shaochilong'') and South America (''Mapusaurus''), though the megaraptorans, which survived until the end of Maastrichtian, may belong to the group as well. A frontal assigned to an allosauroid found to be most closely related to ''Sinraptor'' has also been found in the Coniacian (89-86.3 ...
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Megaraptora
Megaraptora is a clade of carnivorous Tetanurae, tetanuran theropod dinosaurs with controversial relations to other theropods. Its Derived (phylogenetics), derived members, the Megaraptoridae are noted for their elongated hand claws and proportionally large arms, which are usually reduced in size in other large theropods. Megaraptorans are incompletely known, and no complete megaraptoran skeleton has been found. However, they still possessed a number of unique features. Their forelimbs were large and strongly built, and the Glossary of dinosaur anatomy#ulna, ulna bone had a unique shape in members of the Family (biology), family Megaraptoridae, a subset of megaraptorans which excludes ''Fukuiraptor'' and ''Phuwiangvenator''. The first two fingers were elongated, with massive curved claws, while the third finger was small. Megaraptoran skull material is very incomplete, but a juvenile ''Megaraptor'' described in 2014 preserved a portion of the snout, which was long and slender. Le ...
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Orionides
Orionides is a clade of tetanuran theropod dinosaurs from the Middle Jurassic to the Present. The clade includes most theropod dinosaurs, including birds. Relationships Although many phylogenetic analyses found basal tetanurans that were outside both Megalosauroidea and Avetheropoda, the core dichotomy was named only in 2012. Carrano, Benson and Sampson (2012) named that clade Orionides, and defined it as the node comprising Megalosauroidea, Avetheropoda, their most recent common ancestor, and all its descendants. In 2015, Hendrickx, Hartman and Mateus clarified this definition, specifying it as the least inclusive clade including ''Allosaurus fragilis'', ''Megalosaurus bucklandii'', and ''Passer domesticus''. The cladogram presented below follows a phylogenetic analysis published by Zanno and Makovicky in 2013. In 2019, a basal Allosauroid by the name of ''Asfaltovenator'' was discovered that provided more evidence to the idea of a monophyletic Carnosauria that contained both ...
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Monolophosaurus Jiangi
''Monolophosaurus'' ( ; meaning "single-crested lizard") is an extinct genus of tetanuran theropod dinosaur from the Middle Jurassic Shishugou Formation in what is now Xinjiang, China.Holtz, Thomas R. Jr. (2011) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages,'Winter 2010 Appendix./ref> It was named for the single crest on top of its skull. ''Monolophosaurus'' was a mid-sized theropod at about long or more. Discovery and naming A nearly complete skeleton of a theropod new to science was discovered by Dong Zhiming in 1981, during stratigraphic exploration for the benefit of the oil industry. The fossil was not unearthed until 1984. In 1987, before description in the scientific literature, it was referred to in the press as ''Jiangjunmiaosaurus'', an invalid ''nomen nudum''. In 1992 it was mentioned by Dong Zhiming as ''Monolophosaurus jiangjunmiaoi'', and in 1993 by Wayne Grady as ''Monolophosaurus dongi''. These latter names also lacked a ...
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Chuandongocoelurus
''Chuandongocoelurus'' ( ) is a genus of carnivorous tetanuran theropod dinosaur from the Jurassic of China. Discovery and naming The type species ''Chuandongocoelurus primitivus'' was first described and named by Chinese paleontologist He Xinlu in 1984. The generic name combines references to the Chuandong in Sichuan Province and the theropod genus ''Coelurus'', itself named after the Greek κοῖλος, ', meaning "hollow" and οὐρά, ', meaning "tail". The specific name means "the primitive one" in Latin, a reference to the great age of the find. He assigned two partial skeletons to ''Chuandongocoelurus''. The holotype, a thighbone, is part of specimen CCG 20010. Vertebrae, pelvic bones and hindlimb elements, also catalogued under this inventory number, may belong to the same individual. The specimen has unfused neurocentral sutures in its vertebrae, meaning that the animal was immature at the time of death. The second specimen, CCG 20011, is a set of neck vertebrae ...
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Cryolophosaurus
''Cryolophosaurus'' ( or ; "CRY-oh-loaf-oh-SAWR-us") is a genus of large theropod dinosaur known from only a single species ''Cryolophosaurus ellioti'', from the early Jurassic of Antarctica. It was one of the largest theropods of the Early Jurassic, with the subadult being estimated to have reached long and weighed . ''Cryolophosaurus'' was first excavated from Antarctica's Early Jurassic, Pliensbachian aged Hanson Formation, formerly the upper Falla Formation, by paleontologist Dr. William Hammer in 1991. It was the first carnivorous dinosaur to be discovered in Antarctica and the first non-avian dinosaur from the continent to be officially named. The sediments in which its fossils were found have been dated at ~194 to 188 million years ago, representing the Early Jurassic Period. ''Cryolophosaurus'' is known from a skull, a femur and other material, all of which have caused its classification to vary greatly. The femur possesses many primitive characteristics that have cla ...
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Carnosauria
Carnosauria is an extinct large group of predatory dinosaurs that lived during the Jurassic and Cretaceous periods. Starting from the 1990s, scientists have discovered some very large carnosaurs in the carcharodontosaurid family, such as ''Giganotosaurus'', ''Mapusaurus'', ''Carcharodontosaurus'' and ''Tyrannotitan'' which are among the largest known predatory dinosaurs. While it originally contained a wide assortment of giant theropods that were not closely related, the group has since been defined to encompass only the allosaurs and their closest kin. However, with the description and publication in 2019 of ''Asfaltovenator vialidadi'', a basal allosauroid curiously displaying both primitive and derived features seen in Tetanurae, the new phylogenetic analysis has found Megalosauroidea to be a basal grade of carnosaurs in respect to Allosauroidea; thus significantly expanding Carnosauria's inclusiveness towards its original context. Distinctive characteristics of carnosaurs ...
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Monolophosaurus
''Monolophosaurus'' ( ; meaning "single-crested lizard") is an extinct genus of tetanuran theropod dinosaur from the Middle Jurassic Shishugou Formation in what is now Xinjiang, China.Holtz, Thomas R. Jr. (2011) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages,'Winter 2010 Appendix./ref> It was named for the single crest on top of its skull. ''Monolophosaurus'' was a mid-sized theropod at about long or more. Discovery and naming A nearly complete skeleton of a theropod new to science was discovered by Dong Zhiming in 1981, during stratigraphic exploration for the benefit of the oil industry. The fossil was not unearthed until 1984. In 1987, before description in the scientific literature, it was referred to in the press as ''Jiangjunmiaosaurus'', an invalid '' nomen nudum''. In 1992 it was mentioned by Dong Zhiming as ''Monolophosaurus jiangjunmiaoi'', and in 1993 by Wayne Grady as ''Monolophosaurus dongi''. These latter names also lacked a ...
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Sinosaurus
''Sinosaurus'' (meaning "Chinese lizard") is an extinct genus of theropod dinosaur which lived during the Early Jurassic Period. It was a bipedal carnivore approximately in length and in body mass. Fossils of the animal were found at the Lufeng Formation, in the Yunnan Province of China. Discovery and naming The composite term ''Sinosaurus'' comes from ''Sinae'', the Latin word for the Chinese, and the Greek word ' () meaning "lizard"; thus "Chinese lizard". The specific name, ''triassicus'', refers to the Triassic, the period that the fossils were originally thought to date from. ''Sinosaurus'' was described and named by Chung Chien Young, who is known as the 'Father of Chinese Vertebrate Paleontology', in 1940. The holotype, IVPP V34, was found in the Lower Lufeng Formation, and consists of two maxillary (upper jaw) fragments, four maxillary teeth, and a lower jaw fragment with three teeth. The teeth are laterally compressed, and feature fine serrations both at their ant ...
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Theropoda
Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally carnivorous, although a number of theropod groups evolved to become herbivores and omnivores. Theropods first appeared during the Carnian age of the late Triassic Period (geology), period 231.4 million years ago (Year#SI prefix multipliers, Ma) and included all the large terrestrial carnivores from the Early Jurassic until at least the close of the Cretaceous, about 66 Ma. In the Jurassic, birds evolved from small specialized coelurosaurian theropods, and are today represented by about 10,500 living species. Biology Diet and teeth Theropods exhibit a wide range of diets, from insectivores to herbivores and carnivores. Strict carnivory has always been considered the ancestral diet for theropods as a group, and a wider variety of di ...
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Avetheropoda
Avetheropoda, or "bird theropods", is a clade that includes carnosaurians and coelurosaurs to the exclusion of other dinosaurs. Definition Avetheropoda was named by Gregory S. Paul in 1988, and was first defined as a clade by Currie and Padian in 1997, to include ''Allosaurus'', modern birds, and other animals descended from their most recent ancestor. In 1999, Paul Sereno named another group, Neotetanurae, for the clade containing Allosauroidea and Coelurosauria, and excluding other tetanurans such as megalosauroids, but this definition was published slightly later. A monophyletic Avetheropoda is recovered in many papers, however recent findings suggest a monophyletic Carnosauria model with allosauroids and megalosauroids being each other’s closest relatives instead of Allosaurs and Coelurosaurs. Classification The cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladog ...
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Megalosauroidea
Megalosauroidea (meaning 'great/big lizard forms') is a superfamily (or clade) of tetanuran theropod dinosaurs that lived from the Middle Jurassic to the Late Cretaceous period. The group is defined as ''Megalosaurus bucklandii'' and all taxa sharing a more recent common ancestor with it than with ''Allosaurus fragilis'' or ''Passer domesticus''. Members of the group include ''Spinosaurus'', ''Megalosaurus'', and ''Torvosaurus''. They are possibly paraphyletic in nature. Classification The name Spinosauroidea is sometimes used in place of Megalosauroidea. The superfamily Spinosauroidea was named in 1915 by Ernst Stromer. It is a synonym of Megalosauroidea in almost all modern phylogenetic analyses, and it is therefore redundant. Spinosauroidea was defined as a clade in 1998 by Paul Sereno as the node clade containing the common ancestor of ''Spinosaurus'' and ''Torvosaurus'' and all its descendants. Thomas Holtz in 2004 defined a branch clade with the same name containing all spec ...
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