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Orionides
Orionides is a clade of tetanuran theropod dinosaurs from the Middle Jurassic to the Present. The clade includes most theropod dinosaurs, including birds. Relationships Although many phylogenetic analyses found basal tetanurans that were outside both Megalosauroidea and Avetheropoda, the core dichotomy was named only in 2012. Carrano, Benson and Sampson (2012) named that clade Orionides, and defined it as the node comprising Megalosauroidea, Avetheropoda, their most recent common ancestor, and all its descendants. In 2015, Hendrickx, Hartman and Mateus clarified this definition, specifying it as the least inclusive clade including ''Allosaurus fragilis'', ''Megalosaurus bucklandii'', and ''Passer domesticus''. The cladogram presented below follows a phylogenetic analysis published by Zanno and Makovicky in 2013. In 2019, a basal Allosauroid by the name of ''Asfaltovenator'' was discovered that provided more evidence to the idea of a monophyletic Carnosauria that contained both ...
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Megalosauroidea
Megalosauroidea (meaning 'great/big lizard forms') is a superfamily (or clade) of tetanuran theropod dinosaurs that lived from the Middle Jurassic to the Late Cretaceous period. The group is defined as ''Megalosaurus bucklandii'' and all taxa sharing a more recent common ancestor with it than with ''Allosaurus fragilis'' or ''Passer domesticus''. Members of the group include ''Spinosaurus'', ''Megalosaurus'', and ''Torvosaurus''. They are possibly paraphyletic in nature. Classification The name Spinosauroidea is sometimes used in place of Megalosauroidea. The superfamily Spinosauroidea was named in 1915 by Ernst Stromer. It is a synonym of Megalosauroidea in almost all modern phylogenetic analyses, and it is therefore redundant. Spinosauroidea was defined as a clade in 1998 by Paul Sereno as the node clade containing the common ancestor of ''Spinosaurus'' and ''Torvosaurus'' and all its descendants. Thomas Holtz in 2004 defined a branch clade with the same name containing all spec ...
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Avetheropoda
Avetheropoda, or "bird theropods", is a clade that includes carnosaurians and coelurosaurs to the exclusion of other dinosaurs. Definition Avetheropoda was named by Gregory S. Paul in 1988, and was first defined as a clade by Currie and Padian in 1997, to include ''Allosaurus'', modern birds, and other animals descended from their most recent ancestor. In 1999, Paul Sereno named another group, Neotetanurae, for the clade containing Allosauroidea and Coelurosauria, and excluding other tetanurans such as megalosauroids, but this definition was published slightly later. A monophyletic Avetheropoda is recovered in many papers, however recent findings suggest a monophyletic Carnosauria model with allosauroids and megalosauroids being each other’s closest relatives instead of Allosaurs and Coelurosaurs. Classification The cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladog ...
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Sciurumimus
''Sciurumimus'' ("Squirrel-mimic," named for its tail's resemblance to that of the tree squirrel, ''Sciurus'') is an extinct genus of tetanuran theropod from the Late Jurassic of Germany. It is known from a single juvenile specimen representing the type species, ''Sciurumimus albersdoerferi'', which was found in a limestone quarry close to Painten in Lower Bavaria. The specimen was preserved with traces of feather-like filaments. The ''Sciurumimus'' specimen was first announced in 2011 in an informal presentation, but not formally described and named until the following year. Although originally classified as a basal megalosauroid, later phylogenetic analyses dispute this placement, suggesting it was a basal member of the Orionides. Description ''Sciurumimus'' is known from a single holotype fossil that is exceptionally well-preserved, with full skeleton in complete articulation along with fine details of soft tissue. It is comparable in size and proportions to the juvenile coe ...
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Monolophosaurus
''Monolophosaurus'' ( ; meaning "single-crested lizard") is an extinct genus of tetanuran theropod dinosaur from the Middle Jurassic Shishugou Formation in what is now Xinjiang, China.Holtz, Thomas R. Jr. (2011) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages,'Winter 2010 Appendix./ref> It was named for the single crest on top of its skull. ''Monolophosaurus'' was a mid-sized theropod at about long or more. Discovery and naming A nearly complete skeleton of a theropod new to science was discovered by Dong Zhiming in 1981, during stratigraphic exploration for the benefit of the oil industry. The fossil was not unearthed until 1984. In 1987, before description in the scientific literature, it was referred to in the press as ''Jiangjunmiaosaurus'', an invalid '' nomen nudum''. In 1992 it was mentioned by Dong Zhiming as ''Monolophosaurus jiangjunmiaoi'', and in 1993 by Wayne Grady as ''Monolophosaurus dongi''. These latter names also lacked a ...
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Middle Jurassic
The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations containing land animal fossils include the Forest Marble Formation in England, the Kilmaluag Formation in Scotland,British Geological Survey. 2011Stratigraphic framework for the Middle Jurassic strata of Great Britain and the adjoining continental shelf: research report RR/11/06 British Geological Survey, Keyworth, Nottingham. the Daohugou Beds in China, the Itat Formation in Russia, and the Isalo III Formation of western Madagascar. Paleogeography During the Middle Jurassic Epoch, Pangaea began to separate into Laurasia and Gondwana, and the Atlantic Ocean formed. Eastern Laurasia was tectonically active as the Cimmerian plate continued to collide with Laurasia's southern coast, completely closing the Paleo-Tethys Ocean. A subduction zone ...
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Hesperornis Regalis
''Hesperornis'' (meaning "western bird") is a genus of cormorant-like bird that spanned the first half of the Campanian age of the Late Cretaceous period (83.5–78 mya). One of the lesser-known discoveries of the paleontologist O. C. Marsh in the late 19th century Bone Wars, it was an early find in the history of avian paleontology. Locations for ''Hesperornis'' fossils include the Late Cretaceous marine limestones from Kansas and the marine shales from Canada. Nine species are recognised, eight of which have been recovered from rocks in North America and one from Russia. Description ''Hesperornis'' was a large bird, reaching up to in length. It had virtually no wings, and swam with its powerful hind legs. Studies on the feet initially indicated that ''Hesperornis'' and kin had lobed toes similar to modern-day grebes, as opposed to webbed toes as seen in most aquatic birds such as loons. More recent work looking at the morphometrics of the feet in hesperornithiformes and mod ...
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Gregory Scott Paul
Gregory Scott Paul (born December 24, 1954) is an American freelance researcher, author and illustrator who works in paleontology, and more recently has examined sociology and theology. He is best known for his work and research on theropod dinosaurs and his detailed illustrations, both live and skeletal. Professionally investigating and restoring dinosaurs for three decades, Paul received an on-screen credit as dinosaur specialist on ''Jurassic Park'' and Discovery Channel's ''When Dinosaurs Roamed America'' and '' Dinosaur Planet''. He is the author and illustrator of ''Predatory Dinosaurs of the World'' (1988), ''The Complete Illustrated Guide to Dinosaur Skeletons'' (1996), ''Dinosaurs of the Air'' (2001), '' The Princeton Field Guide to Dinosaurs'' (2010), ''Gregory S. Paul's Dinosaur Coffee Table Book'' (2010), ''The Princeton Field Guide to Dinosaurs: 2nd Edition'' (2016), ''The Princeton Field Guide to Pterosaurs'' (2022), ''The Princeton Field Guide to Mesozoic Sea Rep ...
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Coelurosauria
Coelurosauria (; from Greek, meaning "hollow tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyrannosaurs, ornithomimosaurs, and maniraptorans; Maniraptora includes birds, the only known dinosaur group alive today. Most feathered dinosaurs discovered so far have been coelurosaurs. Philip J. Currie had considered it likely and probable that all coelurosaurs were feathered. However, several skin impressions found for some members of this group show pebbly, scaly skin, indicating that feathers did not completely replace scales in all taxa. In the past, Coelurosauria was used to refer to all small theropods, but this classification has since been abolished. Anatomy Bodyplan The studying of anatomical traits in coelurosaurs indicates that the last common ancestor had evolved the ability to eat and digest plant matter, adapting to an omniv ...
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Allosauroidea
Allosauroidea is a superfamily or clade of theropod dinosaurs which contains four family (biology), families — the Metriacanthosauridae, Allosauridae, Carcharodontosauridae, and Neovenatoridae. Allosauroids, alongside the family Megalosauroidea, were among the apex predators that were active during the Middle Jurassic to Late Cretaceous periods. The most famous and best understood allosauroid is the North American genus ''Allosaurus''. The oldest-known allosauroid, ''Shidaisaurus jinae'', appeared in the early Middle Jurassic about 174 million years ago (Earliest Aalenian stage) of China. The last known definitive surviving members of the group died out around 89 million years ago in Asia (''Shaochilong'') and South America (''Mapusaurus''), though the megaraptorans, which survived until the end of Maastrichtian, may belong to the group as well. A frontal assigned to an allosauroid found to be most closely related to ''Sinraptor'' has also been found in the Coniacian (89-86.3 ...
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Carnosauria
Carnosauria is an extinct large group of predatory dinosaurs that lived during the Jurassic and Cretaceous periods. Starting from the 1990s, scientists have discovered some very large carnosaurs in the carcharodontosaurid family, such as ''Giganotosaurus'', ''Mapusaurus'', ''Carcharodontosaurus'' and ''Tyrannotitan'' which are among the largest known predatory dinosaurs. While it originally contained a wide assortment of giant theropods that were not closely related, the group has since been defined to encompass only the allosaurs and their closest kin. However, with the description and publication in 2019 of ''Asfaltovenator vialidadi'', a basal allosauroid curiously displaying both primitive and derived features seen in Tetanurae, the new phylogenetic analysis has found Megalosauroidea to be a basal grade of carnosaurs in respect to Allosauroidea; thus significantly expanding Carnosauria's inclusiveness towards its original context. Distinctive characteristics of carnosaurs ...
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Megaraptora
Megaraptora is a clade of carnivorous Tetanurae, tetanuran theropod dinosaurs with controversial relations to other theropods. Its Derived (phylogenetics), derived members, the Megaraptoridae are noted for their elongated hand claws and proportionally large arms, which are usually reduced in size in other large theropods. Megaraptorans are incompletely known, and no complete megaraptoran skeleton has been found. However, they still possessed a number of unique features. Their forelimbs were large and strongly built, and the Glossary of dinosaur anatomy#ulna, ulna bone had a unique shape in members of the Family (biology), family Megaraptoridae, a subset of megaraptorans which excludes ''Fukuiraptor'' and ''Phuwiangvenator''. The first two fingers were elongated, with massive curved claws, while the third finger was small. Megaraptoran skull material is very incomplete, but a juvenile ''Megaraptor'' described in 2014 preserved a portion of the snout, which was long and slender. Le ...
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Yunyangosaurus
''Yunyangosaurus'' (meaning "Yunyang County lizard") is a genus of possible megalosauroid dinosaur from the Xintiangou Formation in Chongqing, China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and .... The type and only species is ''Yunyangosaurus puanensis''. The name was first published in the 2019 SVP abstract book by Hui (2019) before it was formally described in 2020.Dai Hui, (2019), "A NEW MEGALOSAURID THEROPOD FROM THE MIDDLE JURASSIC XINTIANGOU FORMATION OF CHONGQING, PEOPLE’S REPUBLIC OF CHINA AND ITS IMPLICATION FOR EARLY TETANURAN EVOLUTION", ''SVP 2019, Annual Meeting, Program and Abstracts'', 79A: 86 The specimen consists of a disarticulated partial skeleton consisting of "eleven presacral vertebrae, several cervical and dorsal ribs and chevrons." Judging from the ...
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