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Thelodont
Thelodonti (from Greek: "feeble teeth")Maisey, John G., Craig Chesek, and David Miller. Discovering fossil fishes. New York: Holt, 1996. is a class of extinct jawless fishes with distinctive scales instead of large plates of armor. There is much debate over whether the group of Palaeozoic fish known as the Thelodonti (formerly coelolepids) represent a monophyletic grouping, or disparate stem groups to the major lines of jawless and jawed fish. Thelodonts are united in possession of "thelodont scales". This defining character is not necessarily a result of shared ancestry, as it may have been evolved independently by different groups. Thus the thelodonts are generally thought to represent a polyphyletic group, although there is no firm agreement on this point. On the basis that they are monophyletic, they are reconstructed as being ancestrally marine and invading freshwater on multiple occasions. "Thelodonts" were morphologically very similar, and probably closely related ...
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Thelodont Scales
Thelodonti (from Greek: "feeble teeth")Maisey, John G., Craig Chesek, and David Miller. Discovering fossil fishes. New York: Holt, 1996. is a class of extinct jawless fishes with distinctive scales instead of large plates of armor. There is much debate over whether the group of Palaeozoic fish known as the Thelodonti (formerly coelolepids) represent a monophyletic grouping, or disparate stem groups to the major lines of jawless and jawed fish. Thelodonts are united in possession of "thelodont scales". This defining character is not necessarily a result of shared ancestry, as it may have been evolved independently by different groups. Thus the thelodonts are generally thought to represent a polyphyletic group, although there is no firm agreement on this point. On the basis that they are monophyletic, they are reconstructed as being ancestrally marine and invading freshwater on multiple occasions. "Thelodonts" were morphologically very similar, and probably closely related, ...
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Fish Scale
A fish scale is a small rigid plate that grows out of the skin of a fish. The skin of most jawed fishes is covered with these protective scales, which can also provide effective camouflage through the use of reflection and colouration, as well as possible hydrodynamic advantages. The term ''scale'' derives from the Old French , meaning a shell pod or husk. Scales vary enormously in size, shape, structure, and extent, ranging from strong and rigid armour plates in fishes such as shrimpfishes and boxfishes, to microscopic or absent in fishes such as eels and anglerfishes. The morphology of a scale can be used to identify the species of fish it came from. Scales originated within the jawless ostracoderms, ancestors to all jawed fishes today. Most bony fishes are covered with the cycloid scales of salmon and carp, or the ctenoid scales of perch, or the ganoid scales of sturgeons and gars. Cartilaginous fishes (sharks and rays) are covered with placoid scales. Some sp ...
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Jawless Fish
Agnatha (, Ancient Greek 'without jaws') is an infraphylum of jawless fish in the phylum Chordata, subphylum Vertebrata, consisting of both present (Cyclostomata, cyclostomes) and extinct (conodonts and ostracoderms) species. Among recent animals, cyclostomes are sister taxon, sister to all vertebrates with jaws, known as gnathostomes. Recent molecular data, both from rRNA and from mtDNA as well as embryological data, strongly supports the hypothesis that living agnathans, the cyclostomes, are monophyletic. The oldest fossil agnathans appeared in the Cambrian, and two groups still survive today: the lampreys and the hagfish, comprising about 120 species in total. Hagfish are considered members of the subphylum Vertebrata, because they secondarily lost vertebrae; before this event was inferred from molecular and developmental data, the group Craniata was created by Carl Linnaeus, Linnaeus (and is still sometimes used as a strictly morphological descriptor) to reference hagfi ...
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Agnatha
Agnatha (, Ancient Greek 'without jaws') is an infraphylum of jawless fish in the phylum Chordata, subphylum Vertebrata, consisting of both present ( cyclostomes) and extinct (conodonts and ostracoderms) species. Among recent animals, cyclostomes are sister to all vertebrates with jaws, known as gnathostomes. Recent molecular data, both from rRNA and from mtDNA as well as embryological data, strongly supports the hypothesis that living agnathans, the cyclostomes, are monophyletic. The oldest fossil agnathans appeared in the Cambrian, and two groups still survive today: the lampreys and the hagfish, comprising about 120  species in total. Hagfish are considered members of the subphylum Vertebrata, because they secondarily lost vertebrae; before this event was inferred from molecular and developmental data, the group Craniata was created by Linnaeus (and is still sometimes used as a strictly morphological descriptor) to reference hagfish plus vertebrates. While ...
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Furcacauda
''Furcacauda'' is a genus of thelodontid agnathan Agnatha (, Ancient Greek 'without jaws') is an infraphylum of jawless fish in the phylum Chordata, subphylum Vertebrata, consisting of both present (cyclostomes) and extinct (conodonts and ostracoderms) species. Among recent animals, cyclostomes ... from the Lower Devonian of Canada, and is the type genus of the order Furcacaudiformes. Furcacaudiform thelodontids were deep water jawless vertebrates with symmetrical fork and lobed-finned tails and scales smaller than typical loganellid and nikoliviid thelodonti scales. Furcacaudiform thelodonts are noted as having a laterally compressed body, large anterior eyes, slightly posterior, lateral, and vertical to a small mouth, and a condensed curved row of branchial openings (gills) directly posterior to the eyes. Many but not all had laterally paired fins. Wilson and Caldwell also note the presence of a caudal peduncle and a long caudal fin made of two large lobes, one dorsal and ...
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Archipelepidiformes
Archipelepidiformes is an order of extinct jawless fishes in the class Thelodonti. Archipelepidiforms are regarded as the basalmost thelodonts primarily because the histology and morphology of archipelepidiforms have many similarities with the histology and morphology of pteraspidomorphs, hinting that the two groups share a common ancestor, and hinting that archipelepidiforms retain many primitive features from this common ancestor.Soehn, K. L., Märss, T., Caldwell, M. W. & Wilson, M. V. H., 2001: New and biostratigraphically useful thelodonts from the Silurian of the Mackenzie Mountains, Northwest Territories, Canada. Journal of Vertebrate Paleontology, 21: 651-65/ref> Wilson, Mark VH, and Tiiu Märss. "Thelodont phylogeny revisited, with inclusion of key scale-based taxa." Estonian Journal of Earth Sciences 58.4 (2009): 297œ310/ref> Currently, only whole body fossils of '' Archipelepis'' are known: these fossils show that archipelepids were tadpole-like animals with no fins ...
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Furcacaudiformes
Furcacaudiformes is an extinct order of jawless fish in the class Thelodonti. Because the paucity of intact fossils, especially since some families are known entirely from scale fossils, taxonomy of thelodonts is based primarily on scale morphology. A 2009 assessment of thelodont taxonomy by Wilson and Märss merges the orders Loganelliiformes, Katoporiida and Shieliiformes into Thelodontiformes, places families Lanarkiidae and Nikoliviidae into Furcacaudiformes (because of scale morphology) and establishes Archipelepidiformes Archipelepidiformes is an order of extinct jawless fishes in the class Thelodonti. Archipelepidiforms are regarded as the basalmost thelodonts primarily because the histology and morphology of archipelepidiforms have many similarities with the h ... as the basal-most order. References External links * Thelodonti Devonian jawless fish Prehistoric jawless fish orders {{Devonian-jawless-fish-stub ...
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Thelodontiformes
Thelodontiformes is an extinct order of jawless fish of the Silurian.Kiaer, J., 1932: New coelolepids from the Upper Silurian on Oesel (Esthonia). Eesti Loodusteaduse Arhiiv, Seeria 1, 10: 167-176. Because the paucity of intact fossils, especially since some families are known entirely from scale fossils, taxonomy of thelodonts is based primarily on scale morphology. A recent assessment of thelodont taxonomy by Wilson and Märss in 2009 merges the orders Loganelliiformes, Katoporiida and Shieliiformes into Thelodontiformes, places families Lanarkiidae and Nikoliviidae into Furcacaudiformes (because of scale morphology) and establishes Archipelepidiformes as the basal-most order.Wilson, Mark VH, and Tiiu Märss. "Thelodont phylogeny revisited, with inclusion of key scale-based taxa." Estonian Journal of Earth Sciences 58.4 (2009): 297œ310/ref> References External links * Thelodonti Silurian jawless fish Prehistoric jawless fish orders Silurian first appeara ...
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Gnathostome
Gnathostomata (; from Greek: (') "jaw" + (') "mouth") are the jawed vertebrates. Gnathostome diversity comprises roughly 60,000 species, which accounts for 99% of all living vertebrates, including humans. In addition to opposing jaws, living gnathostomes have true teeth (a characteristic which has subsequently been lost in some), paired appendages (pectoral and pelvic fins, arms, legs, wings, etc.), the elastomeric protein of elastin, and a horizontal semicircular canal of the inner ear, along with physiological and cellular anatomical characters such as the myelin sheaths of neurons, and an adaptive immune system that has the discrete lymphoid organs of spleen and thymus, and uses V(D)J recombination to create antigen recognition sites, rather than using genetic recombination in the variable lymphocyte receptor gene. It is now assumed that Gnathostomata evolved from ancestors that already possessed a pair of both pectoral and pelvic fins. Until recently these ancestors, know ...
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Lanarkia
''Lanarkia'' is a genus of extinct thelodont agnathan which existed in what is now Scotland and Canada during the upper Silurian The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago (Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoi ... period.''Lanarkia horrida''
at www.fossilmuseum.net.


References

Thelodonti genera Silurian jawless fish
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Late Devonian Extinction
The Late Devonian extinction consisted of several extinction events in the Late Devonian Epoch (geology), Epoch, which collectively represent one of the five largest extinction event, mass extinction events in the history of life on Earth. The term primarily refers to a major extinction, the Kellwasser event (also known as the Frasnian-Famennian extinction), which occurred around 372 million years ago, at the boundary between the Frasnian stage and the Famennian stage, the last Stage (stratigraphy), stage in the Devonian Period.Racki, 2005McGhee, George R. Jr, 1996. The Late Devonian Mass Extinction: the Frasnian/Famennian Crisis (Columbia University Press) Overall, 19% of all families and 50% of all genera became extinct. A second mass extinction, the Hangenberg event (also known as the end-Devonian extinction), occurred 359 million years ago, bringing an end to the Famennian and Devonian, as the world transitioned into the Carboniferous, Carboniferous Period. Although it is we ...
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Morphology (biology)
Morphology is a branch of biology dealing with the study of the form and structure of organisms and their specific structural features. This includes aspects of the outward appearance ( shape, structure, colour, pattern, size), i.e. external morphology (or eidonomy), as well as the form and structure of the internal parts like bones and organs, i.e. internal morphology (or anatomy). This is in contrast to physiology, which deals primarily with function. Morphology is a branch of life science dealing with the study of gross structure of an organism or taxon and its component parts. History The etymology of the word "morphology" is from the Ancient Greek (), meaning "form", and (), meaning "word, study, research". While the concept of form in biology, opposed to function, dates back to Aristotle (see Aristotle's biology), the field of morphology was developed by Johann Wolfgang von Goethe (1790) and independently by the German anatomist and physiologist Karl Fr ...
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