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Tarphycerida
The Tarphycerida were the first of the coiled cephalopods, found in marine sediments from the Lower Ordovician (middle and upper Canad) to the Middle Devonian. Some, such as '' Aphetoceras'' and '' Estonioceras'', are loosely coiled and gyroconic; others, such as '' Campbelloceras'', '' Tarphyceras'', and '' Trocholites'', are tightly coiled, but evolute with all whorls showing. The body chamber of tarphycerids is typically long and tubular,Furnish and Glenister 1964; Nautiloidea - Tarphycerida; In the ''Treatise on Invertebrate Paleontology'' Vol K; Teichert and Moore, (eds) GSA and U of Kansas Press 1964 as much as half the length of the containing whorl in most, greater than in the Silurian Ophidioceratidae. The Tarphycerida evolved from the elongated, compressed, exogastric Bassleroceratidae, probably ''Bassleroceras'', around the end of the Gasconadian through forms like ''Aphetoceras''. Close coiling developed rather quickly, and both gyroconic and evolute forms are fou ...
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Cephalopods
A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles (muscular hydrostats) modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology. Cephalopods became dominant during the Ordovician period, represented by primitive nautiloids. The class now contains two, only distantly related, extant subclasses: Coleoidea, which includes octopuses, squid, and cuttlefish; and Nautiloidea, represented by ''Nautilus'' and ''Allonautilus''. In the Coleoidea, the molluscan shell has been internalized or is absent, whereas in the Nautiloidea, the external shell remains. About 800 living species of cephalopods have been identified. Tw ...
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Tarphyceratidae
The Tarphyceratidae are tightly coiled, evolute Tarphycerida with ventral siphuncles. The dorsum is characteristically impressed where the whorl presses against the venter of the previous. The Tarphyceratidae are derived from ''Bassleroceras'' or possibly from some member of the Estonioceratidae. Tarphyceratids vary in form and siphuncle position. The siphuncle may be subcentral as in ''Centrotarphyceras''; intermediary as in '' Tarphyceras'', or subventral as in '' Campbelloceras''. Cross sections vary. ''Tarphyceras'' and ''Campbelloceras'' are rounded. ''Centrotarphyceras'' is subquadrate with broadly rounded flanks and a slightly rounded venter. ''Eurystomites'' is subquadrate and slightly compressed, with the maximum width closer to the dorsum. ''Pionoceras'' is subrounded and slightly depressed with the maximum width closer to the venter. The dorsal impression in ''Tarphyceras'' and ''Eurystomites'' is broad and deep and in ''Pionoceras'' and ''Centrotarphyceras'', broad an ...
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Bassleroceras
''Bassleroceras'' is an elongate upwardly curved, exogastric, genus with the venter on the under side more sharply rounded than the dorsum on the upper. The siphuncle is ventral, composed of thick-walled tubular segments in which connection rings thicken in towardly as in both the Ellesmerocerida and primitive Tarphycerida. ''Bassleroceras'' is the type genus of the Bassleroceratidae which Furnish and Glenister (1964) included in the Ellesmerocerida, but which Flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ... (1967) placed in the Tarphycerida. ''Bassleroceras'' gave rise to the Tarphycerida (sensu Furnish and Glenister, 1964) by evolving genera with tighter and tighter curvatures until becoming gyroconic, a character of the Estonioceratidae, a family of early tarphycerids ...
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Bassleroceratidae
The Bassleroceratidae is a family of gradually expanding, smooth ellesmerocerids with a slight to moderate exogastric curvature, subcircular to strongly compressed cross section, and ventral orthochaonitc siphuncle. The ventral side is typically more sharply rounded than the dorsal side and septa are close spaced. Connecting rings are thick and slightly expanded into the siphuncle, making the segments slightly concave; characteristic of the Ellesmerocerida.Furnish and Glenister 1964a, Nautiloidea-Ellesmerocerida; Treatise on Invertebrate Paleontology, Part K Nautiloidea. Basslerocerids are limited to the Lower Ordovician and first appeared sometime in the Gasconadian, (Tremedocian) They gave rise, possibly through ''Bassleroceras'', by evolving and ever tightened curvature to the Tarphycerida and by a thinning of the connecting rings to the Graciloceratidae which are ancestral Oncocerida The Oncocerida comprise a diverse group of generally small nautiloid cephalopods known fr ...
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Aphetoceras
''Aphetoceras'' is a genus of tarphycerid cephalopod within the Estonioceratidae; loosely coiled without an impression along the dorsal margin; early whorls barely reaching, separating then diverging in the final mature whorl; weakly ribbed in some. The cross section of ''Aphetoceras'' is higher than wide, making it compressed in form. The dorsum, along the inner curve, is more broadly rounded than the venter which lies along the outer curve. The siphuncle is relatively large, located near but not at the ventral margin; lined with secondary deposits. Chambers are empty. ''Aphetoceras'' is similar in outline to the related '' Estonioceras'' and ''Alaskoceras'' but most similar in cross section to '' Clytoceras''. ''Aphetoceras'' comes from the Lower Ordovician of North America and Australia and is one of the earlier tarphycerids found in the Lower Ordovician The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geo ...
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Ophidioceratidae
''Ophioceras'' is a genus of closely coiled tarphycerid nautiloid cephalopods, the sole representatives of the family Ophidioceratidae, characterized by an evolute shell with narrow, subrounded, annulated whorls and a subcentral siphuncle composed of thin connecting rings that show no evidence of layering. The mature body chamber is strongly divergent and is the longest proportionally of any tarphycerid. The aperture has a deep hyponomic sinus and ocular sinuses, and so resembles some lituitids. ''Ophioceras'' seem to have its origin in the Trocholitidae, possibly in '' Graftonoceras''. W.M Furnish and Brian F Glenister 1964. Nautiloidea-Tarphyhceria. Treatise on Invertebrate Paleontology Part K. Geological Soc of America and Univ Kansas Press. ''Ophioceras'' has a narrow, evolutely wound, closely coiled shell that diverges in the adult stage. The shell is more or less circular in cross section, strongly annulate with close spaced, rounded, transverse ribs. The venter is b ...
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Trocholitidae
The Trocholitidae are Tarphycerida with whorls in close contact as with the Tarphyceratidae, but in which the siphuncle, similar in structure, becomes dorsal. The Trocholitidae are derived from the Tarphyceratidae, perhaps from different tarphyceratids. Members of the Trocholitidae vary in whorl section and siphuncle position. Most are tightly coiled with the dorsum impressed to some degree. As exceptions ''Arkoceras'' and ''Wichitoceras'' have their whorls touching but without a dorsal impression. '' Discoceras'' and '' Curtoceras'' have somewhat subquadrate whorl sections and are moderately impressed. In ''Graftonoceras'' and ''Jasperoceras'' the venter is rounded and continues smoothly through the flanks to a broad dorsum which in each is moderately impressed. The siphuncle in ''Graftonoceras'' is on the dorsal margin, in ''Jasperoceras'' it is between the center and dorsum. Whorls in T''rocholites'' have a wide cross section and low profile, in ''Wichitoceras'' they are lat ...
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Tarphyceras
''Tarphyeras'' is a genus of tarphyceratid with whorls rounded in cross section, having a deeply impressed dorsum and a ventral to subcentral siphuncle, known from the Lower Ord (U Canad) of North America. It differs from '' Campbelloceras'' in that ''Campbelloceras'' is only slightly impressed, from '' Centrotarphyceras'' in that ''Centrotarphyceras'' is subquadrate and has a central siphuncle, and from ''Trocholites'' in that although ''Trocholites'' is subcircular in cross section, the siphuncle is subdorsal. ''Taphyceras'' may be derived from ''Campbelloceras'' through development of tighter coiling and a deeper impression along with a smaller siphuncle The siphuncle is a strand of tissue passing longitudinally through the shell of a cephalopod mollusk. Only cephalopods with chambered shells have siphuncles, such as the extinct ammonites and belemnites, and the living nautiluses, cuttlefish, and ... that may be further offset from the venter. ''Tarphyceras'' was named by ...
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Estonioceratidae
The Estonioceratidae are a family of loosely coiled tarphycerids in which the inner side of the whorls, which forms the dorsum, is rounded or flat with no impression, and in which the siphuncle, composed of thick tubular segments, is located ventrally. The Estonioceratidae seem to form a link between the ancestral Bassleroceratidae and the more tightly coiled Tarphyceratidae The Tarphyceratidae are tightly coiled, evolute Tarphycerida with ventral siphuncles. The dorsum is characteristically impressed where the whorl presses against the venter of the previous. The Tarphyceratidae are derived from ''Bassleroceras'' or ....Furnish, W. M, and Glenister, Brian F. 1964. Nautiloidea – Tarphycerida; Treatise on Invertebrate Paleontology, Part K, Geol. Soc. of America, Teichert and Moore (eds.) References {{Taxonbar, from=Q16965822 Cephalopod families Nautiloids Ordovician cephalopods ...
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Trocholites
''Trocholites'' is a tarphycerid genus in the family Trocholitidae from the Middle and Late Ordovician with a gradually expanding, weakly ribbed shell; whorls in contact, dorsum slightly impressed; cross section depressed, venter and sides rounded; siphuncle close to but not at the dorsal margin. The dorsal siphuncle of ''Trocholites'', and the Trocholitidae is somewhat anomalous and may indicate something about the orientation of the shell during life; placing the last septum high rather than midway or low, typical of most. The prevailing location in most coiled cephalopods is central or ventral, extremely ventral in most ammonoids. '' Trochoceras'', of similar name, is a member of the Rutoceratidae in the Nautilida, and therefore only very distantly related back through the Bassleroceratidae The Bassleroceratidae is a family of gradually expanding, smooth ellesmerocerids with a slight to moderate exogastric curvature, subcircular to strongly compressed cross section, and ven ...
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Barrandeoceratidae
The Barrandeoceratidae is a family of coiled nautiloids included in the Tarphycerida (''sensu lato'') that lived from the Middle Ordovician to the Middle Devonian, characterised by mostly compressed shells with a subcentral siphuncle composed of thin-walled segments that may become secondarily ventral.(Flower and Kummel 1950, Sweet 1964). The Barrandeoceratidae are derived from the tarphyceratid genus, ''Centrotarphyceras'' through '' Barrandeoceras'' (Flower 1984), and is the source for the Uranoceratidae and Nephriticeratidae. These three families form the barrandeoceratid group. Sometimes a fourth family, the Bickmoritidae, is added, based on the genus ''Bickmorites''. Genera (Flower 1984) include '' Barrandeoceras'', '' Centrocyrtoceras'', '' Paquettoceras'', '' Savageoceras'', '' Gasconsoceras'', ''Haydenoceras'', and '' Paraplectoceras''; and if included, '' Bickmorites''. ''Avilionella'' and ''Laureloceras'', included in the Barrandeocerida in the ''Treatise on Invert ...
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Campbelloceras
''Campbelloceras'' is a tarphyceratid nautiloid known from the Lower Ordovician, Upper Canadian Epoch of North America, where it is widespread. ''Campbelloceras'' was named by Ulrich and Foerste in 1936. The shell of ''Campbelloceras'' has a circular whorl section, only slightly impressed, and a siphuncle that is close to the venter in all growth stages. ''Campbelloceras'' differs from ''Tarphyceras'' in that the rate of expansion is greater, the siphuncle is proportionally largers, and an impression is shallower. ''Campbelloceras'' may have given rise to the barrandeocerid '' Plectoceras'' (Plectoceratidae) through simplification of the connecting rings, and to ''Tarphyceras'' through tighter coiling, development of a deeper impression, and reduction in the size of the siphuncle. ''Campbelloceras'' may be conceivably derived from '' Estonioceras'' (Estonioceratidae). The cross section of ''Campbelloceras'' and that of the outer whorl of ''Estonioceras'' are similar, except f ...
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