Tarphyceratidae
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The Tarphyceratidae are tightly coiled, evolute Tarphycerida with ventral
siphuncle The siphuncle is a strand of tissue passing longitudinally through the shell of a cephalopod mollusk. Only cephalopods with chambered shells have siphuncles, such as the extinct ammonites and belemnites, and the living nautiluses, cuttlefish, and ...
s. The dorsum is characteristically impressed where the whorl presses against the venter of the previous. The Tarphyceratidae are derived from ''
Bassleroceras ''Bassleroceras'' is an elongate upwardly curved, exogastric, genus with the venter on the under side more sharply rounded than the dorsum on the upper. The siphuncle is ventral, composed of thick-walled tubular segments in which connection ring ...
'' or possibly from some member of the Estonioceratidae. Tarphyceratids vary in form and siphuncle position. The siphuncle may be subcentral as in ''Centrotarphyceras''; intermediary as in '' Tarphyceras'', or subventral as in '' Campbelloceras''. Cross sections vary. ''Tarphyceras'' and ''Campbelloceras'' are rounded. ''Centrotarphyceras'' is subquadrate with broadly rounded flanks and a slightly rounded venter. ''Eurystomites'' is subquadrate and slightly compressed, with the maximum width closer to the dorsum. ''Pionoceras'' is subrounded and slightly depressed with the maximum width closer to the venter. The dorsal impression in ''Tarphyceras'' and ''Eurystomites'' is broad and deep and in ''Pionoceras'' and ''Centrotarphyceras'', broad and shallow. In ''Campbelloceras'' and ''Cycloplectoceras'' the impression is slight, running along the dorsal median. The Tarphyceratidae are known mainly from the
Lower Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. The ...
(M-U Canad); gave rise to the
Trocholitidae The Trocholitidae are Tarphycerida with whorls in close contact as with the Tarphyceratidae, but in which the siphuncle, similar in structure, becomes dorsal. The Trocholitidae are derived from the Tarphyceratidae, perhaps from different tarphyc ...
and provided the sources for the more advanced barrandeocerids


Genera

Genera in the Tarphyeratidae as named in the
Treatise on Invertebrate Paleontology The ''Treatise on Invertebrate Paleontology'' (or ''TIP'') published by the Geological Society of America and the University of Kansas Press, is a definitive multi-authored work of some 50 volumes, written by more than 300 paleontologists, and co ...
Part K (Furnish and Glenister 1964) are restricted to the middle and upper parts of the Lower Ordovician in North America. For purposes of discussion only, they are divided into those where dorsum, which is on the inside or concave side of the volutions, is more deeply impressed and those with dorsum that is only moderately or slightly impressed. (Fig 262,K359,Furnish and Glenister, 1964) Tarphyceratidae with dorsum deeply impressed * '' Tarphyceras'':serpenticonic, whorls rounded vantrally and laterally, deeply impressed dorsally; rate of expansion moderate; siphuncle ventral to subcentral. Found widespread. * '' Centrotarphyceras'': Whorl section generally subquadrate, typically with flattened venter and flanks. Expanded more gradually than Tarphyceras. Mature siphuncle subcenral. Widespread * '' Eurystomites'': Large, whorls subquadrate, may be higher than wide, broadly rounded ventrally and laterally with discernible ventrolateral shoulders, dorsally impressed; siphuncle large, subcentral Widespread. * '' Pionoceras'' : Whorl section suboval with broadly rounded venter and flanks that converge on the more narrow and impressed dorsum. Siphuncle fairly large, submarginal. Widespread Tarphycerida with dorsum slightly to moderately impressed * '' Campbelloceras'': whorl section circular, slightly impressed; siphuncle close to the venter in all growth stages. Expanded more rapidly than Tarphyceras, siphuncle proportionally larger. Widespread. * '' Cycloplectoceras'': like ''Campbelloceras'' but with pronounced ribs. Found in Arkansas. * '' Moreauoceras'': like ''campbelloceras'' but more gradually expanded and with subterminal mature aperture. Found in Missouri * '' Pilotoceras'': small, whorls subtapezoidal to subtriangular in section; venter narrow, sides flattened and divergent, dorsum with small shallow impression; siphuncle narrow, subcentral. Widespread. Uncertain *'' Seelyoceras'': Depressed, strongly annulate (transversely ribbed), curved nautiloid with ventral marginal siphuncle based on a single fragments. Affinities uncertain. Found in New York (state).


References

*Flower R.H. 1976; Ordovician Cephalopod Faunas and Their Role in Correlation; pp523–552 in Basset, M.G. (Ed) 1976. The Ordovician System: proceedings of a Palaeontological Association symposium, 1964... University of Wales press and National Museum of Wales, Cardiff. *Furnish, W.M. and Glenister, Brian F 1964; Nautiloidea -Tarphycerida, in The Treatise on Invertebrate Paleontology, Part K, Nautiloidea; Geological Society of America and University of Kansas press. {{Taxonbar, from=Q7686585 Tarphycerida Cephalopod families Early Ordovician first appearances Early Ordovician extinctions