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Tapejarid
Tapejaridae (from a Tupi word meaning "the old being") are a family of pterodactyloid pterosaurs from the Cretaceous period. Members are currently known from Brazil, England, Hungary, Morocco, Spain, the United States, and China. The most primitive genera were found in China, indicating that the family has an Asian origin. Description Tapejarids were small to medium-sized pterosaurs with several unique, shared characteristics, mainly relating to the skull. Most tapejarids possessed a bony crest arising from the snout (formed mostly by the premaxillary bones of the upper jaw tip). In some species, this bony crest is known to have supported an even larger crest of softer, fibrous tissue that extends back along the skull. Tapejarids are also characterized by their large nasoantorbital fenestra, the main opening in the skull in front of the eyes, which spans at least half the length of the entire skull in this family. Their eye sockets were small and pear-shaped. Studies of tapejarid ...
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Sinopterus Dongi
''Sinopterus'' (meaning "Chinese wing") was a genus of tapejarid pterodactyloid pterosaur from the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. It was first described and named by Wang Xiaolin and Zhou Zhonghe. Three species have been classified in this genus, though only two are generally considered to be valid. ''Sinopterus'' is known for its proportionally large skull, which has a birdlike pointed beak, a long bony crest that starts with a tall premaxilla and goes back along the middle of the skull to form a point overhanging the rear of the skull, and its lack of teeth. Description The type species, ''S. dongi'', is based on IVPP V13363, an articulated, nearly complete skeleton. The skull of this individual was 17 centimeters (6.7 inches) long, and the wingspan was estimated to be 1.2 meters (3.9 feet). The authors suggested that it was an omnivore, and noted that it was the first record of a tapejarid outside of Brazil, and the earlies ...
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Sinopterus
''Sinopterus'' (meaning "Chinese wing") was a genus of tapejarid pterodactyloid pterosaur from the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. It was first described and named by Wang Xiaolin and Zhou Zhonghe. Three species have been classified in this genus, though only two are generally considered to be valid. ''Sinopterus'' is known for its proportionally large skull, which has a birdlike pointed beak, a long bony crest that starts with a tall premaxilla and goes back along the middle of the skull to form a point overhanging the rear of the skull, and its lack of teeth. Description The type species, ''S. dongi'', is based on IVPP V13363, an articulated, nearly complete skeleton. The skull of this individual was 17 centimeters (6.7 inches) long, and the wingspan was estimated to be 1.2 meters (3.9 feet). The authors suggested that it was an omnivore, and noted that it was the first record of a tapejarid outside of Brazil, and the earlies ...
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Thalassodrominae
Thalassodrominae or Thalassodromidae (meaning "sea runners", due to previous misconceptions of skimming behavior; they are now thought to be terrestrial predators) is a group of azhdarchoid pterosaurs from the Cretaceous period. Its traditional members come from Brazil, however, other possible members also come from other places, including the United States, Morocco, and Argentina. Thalassodrominae is considered either to be a subfamily within the pterosaur family Tapejaridae, or as a distinct family, Thalassodromidae, within the clade Neoazhdarchia, closely related to dsungaripterids or azhdarchids. Classification Thalassodrominae traditionally includes only two genera, ''Thalassodromeus'' and ''Tupuxuara'', and was defined to include them and any other descendants of their most recent common ancestor. The classification of thalassodromines is controversial. Some studies, including one by Lü and colleagues in 2008, have found that the thalassodromines are more closely related t ...
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Bakonydraco
''Bakonydraco'' is a genus of pterodactyloid pterosaur from the Late Cretaceous period (Santonian stage) of what is now the Csehbánya Formation of the Bakony Mountains, Iharkút, Veszprém, western Hungary. Etymology ''Bakonydraco'' was named in 2005 by paleontologists Attila Ősi, David Weishampel, and Jianu Coralia. The type species is ''Bakonydraco galaczi''. The genus name refers to the Bakony Mountains and to Latin ''draco'', "dragon". The specific epithet ''galaczi'' honors Professor András Galácz, who helped the authors in the Iharkút Research Program, where fossils are since 2000 found in open-pit mining of bauxite, among them the remains of pterosaurs, the first ever discovered in Hungary. Description ''Bakonydraco'' is based on holotype MTM Gyn/3, a nearly complete mandibula, a fusion of the lower jaws. Also assigned to it, as paratype, is MTM Gyn/4, 21: parts from another jaw's symphysis (the front parts, having fused into a single blade-like structure, of th ...
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Huaxiapterus
''Huaxiapterus'' (meaning "Hua Xia hinawing") is a genus of tapejarid pterodactyloid pterosaur from the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. It is the second genus of tapejarid from this formation, after ''Sinopterus''. Three species are known (''H. jii'', "''H. corollatus''", and "''H. benxiensis''"), though they may not actually form a natural group with each other. It was first named by Lü Junchang and Yuan Chongxi. Discovery The type species of ''Huaxiapterus'' is ''H. jii'', which is based on GMN-03-11-001, a nearly complete skeleton and skull. While it was initially separated from the closely related ''Sinopterus'' by several characters including its larger size and larger crest, later studies showed that it was in fact more closely related to ''Sinopterus dongi'' than to either of the other two species of ''Huaxiapterus''. Some researchers have considered ''H. jii'' to be a species of ''Sinopterus'' for this reason, though a mo ...
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Pterosaurs
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or 'rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were insectiv ...
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Tupandactylus
''Tupandactylus'' (meaning "Tupan finger", in reference to the Tupi thunder god) is a genus of tapejarid pterodactyloid pterosaur from the Early Cretaceous Crato Formation of Brazil. History ''Tupandactylus imperator'' is known from four nearly complete skulls. The holotype specimen is MCT 1622-R, a skull and partial lower jaw, found in the Crato Formation, dating to the boundary of the Aptian-Albian stages of the early Cretaceous period, about 112 Ma ago.Martill, D.M., Bechly, G. and Loveridge, R.F. (2007). ''The Crato fossil beds of Brazil: window into an ancient world.'' Cambridge University Press. , It was initially described as a species of '' Tapejara'', but later research has indicated it warrants its own genus. The skull was toothless and had a prominent sagittal crest, only the base of which was bony: the front of the crest featured a tall bony rod extending up and back, and the rear of the crest had a long prong of bone projecting behind it. The bulk of the crest was ...
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Tupandactylus Imperator
''Tupandactylus'' (meaning "Tupan finger", in reference to the Tupi people, Tupi thunder god) is a genus of tapejaridae, tapejarid pterodactyloid pterosaur from the Early Cretaceous Crato Formation of Brazil. History ''Tupandactylus imperator'' is known from four nearly complete skulls. The holotype specimen is Museu de Ciências da Terra, MCT 1622-R, a skull and partial lower jaw, found in the Crato Formation, dating to the boundary of the Aptian-Albian stages of the early Cretaceous period, about 112 Ma ago.Martill, D.M., Bechly, G. and Loveridge, R.F. (2007). ''The Crato fossil beds of Brazil: window into an ancient world.'' Cambridge University Press. , It was initially described as a species of ''Tapejara (pterosaur), Tapejara'', but later research has indicated it warrants its own genus. The skull was toothless and had a prominent sagittal crest, only the base of which was bony: the front of the crest featured a tall bony rod extending up and back, and the rear of the cre ...
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Tapejara Wellnhoferi
''Tapejara'' (from a Tupi word meaning "the old being") is a genus of Brazilian pterosaur from the Cretaceous Period (Santana Formation, dating to about 127 to 112 million years ago). ''Tapejara'' crests consisted of a semicircular crest over the snout, and a bony prong which extended back behind the head. It was a small pterosaur, with a wingspan of approximately . Species and classification The type species and only one currently recognized as valid by most researchers, is ''T. wellnhoferi''. The specific name honors German paleontologist Peter Wellnhofer. Two larger species, originally named ''Tapejara imperator'' and ''Tapejara navigans'', were later classified in the genus ''Tapejara''. However, several studies have shown that ''T. imperator'' and ''T. navigans'' are significantly different from ''T. wellnhoferi'' and therefore were reclassified into new genera. The species ''T. imperator'' was given its own genus, ''Tupandactylus'', by Alexander Kellner and Diogenes de Al ...
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Tapejara (pterosaur)
''Tapejara'' (from a Tupi word meaning "the old being") is a genus of Brazilian pterosaur from the Cretaceous Period (Santana Formation, dating to about 127 to 112 million years ago). ''Tapejara'' crests consisted of a semicircular crest over the snout, and a bony prong which extended back behind the head. It was a small pterosaur, with a wingspan of approximately . Species and classification The type species and only one currently recognized as valid by most researchers, is ''T. wellnhoferi''. The specific name honors German paleontologist Peter Wellnhofer. Two larger species, originally named ''Tapejara imperator'' and ''Tapejara navigans'', were later classified in the genus ''Tapejara''. However, several studies have shown that ''T. imperator'' and ''T. navigans'' are significantly different from ''T. wellnhoferi'' and therefore were reclassified into new genera. The species ''T. imperator'' was given its own genus, ''Tupandactylus'', by Alexander Kellner and Diogenes de A ...
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Eopteranodon
''Eopteranodon'' (meaning "dawn '' Pteranodon'' (toothless wing)") is a genus of tapejarid pterosaur from the Aptian-age Lower Cretaceous Yixian Formation of Beipiao City, Liaoning, China. The genus was named in 2005 by paleontologists Lü Junchang and Zhang Xingliao. The type and only species is ''Eopteranodon lii''. Description and classification ''Eopteranodon'' is based on the type specimen or holotype BPV-078, an incomplete skeleton and skull. Its skull, including a large crest, was toothless and similar to that of ''Pteranodon''. The skull lacks the point of the snout but it was in life less than 200 millimeters long (7.9 inches), and the animal had a wingspan of about 1.1 meters (3.6 feet). A second specimen, D2526, described in 2006, had a larger wingspan. Despite its similarities to ''Pteranodon'', ''Eopteranodon'' was not placed into a family by its describers, who put it into the clade Pteranodontia as ''incertae sedis'' (uncertain position). Sho ...
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Aymberedactylus
''Aymberedactylus'' (meaning "small lizard finger") is a genus of tapejarid pterosaur from the Early Cretaceous Crato Formation of Brazil. It contains a single species, ''A. cearensis.'' Discovery and naming The holotype specimen of ''Aymberedactylus'', MN 7596-V, is a nearly complete mandible preserved in three dimensions. It was discovered in the Aptian- Albian Crato Formation, a konservat-lagerstätte well known for its exquisite preservation of fossils, and it was described in 2016. The name ''Aymberedactylus'' is derived from the Tupi word ("small lizard") and Greek ("finger"), and the specific name refers to the Brazilian state of Ceará, which it was discovered in. Description ''Aymberedactylus'' can be identified as a tapejarine tapejarid from the holotypic jawbone due to its toothlessness, slightly downturned dentary symphysis which accounts for half of the total length of the jaw, and a small crest on the bottom of the dentary (which was incompletely preserved ...
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