Symphyta
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Symphyta
Sawflies are the insects of the suborder Symphyta within the order Hymenoptera, alongside ants, bees, and wasps. The common name comes from the saw-like appearance of the ovipositor, which the females use to cut into the plants where they lay their eggs. The name is associated especially with the Tenthredinoidea, by far the largest superfamily in the suborder, with about 7,000 known species; in the entire suborder, there are 8,000 described species in more than 800 genera. Symphyta is paraphyletic, consisting of several basal groups within the order Hymenoptera, each one rooted inside the previous group, ending with the Apocrita which are not sawflies. The primary distinction between sawflies and the Apocrita – the ants, bees, and wasps – is that the adults lack a "wasp waist", and instead have a broad connection between the abdomen and the thorax. Some sawflies are Batesian mimics of wasps and bees, and the ovipositor can be mistaken for a stinger. Sawflies vary in leng ...
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Sawfly (Nematus Miliaris) Larvae
Sawflies are the insects of the suborder Symphyta within the order Hymenoptera, alongside ants, bees, and wasps. The common name comes from the saw-like appearance of the ovipositor, which the females use to cut into the plants where they lay their eggs. The name is associated especially with the Tenthredinoidea, by far the largest Taxonomic rank#Ranks in zoology, superfamily in the suborder, with about 7,000 known species; in the entire suborder, there are 8,000 described species in more than 800 genera. Symphyta is paraphyly, paraphyletic, consisting of several Basal (phylogenetics), basal groups within the order Hymenoptera, each one rooted inside the previous group, ending with the Apocrita which are not sawflies. The primary distinction between sawflies and the Apocrita – the ants, bees, and wasps – is that the adults lack a "wasp waist", and instead have a broad connection between the abdomen and the Thorax (insect anatomy), thorax. Some sawflies are Batesian mimicry, ...
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Hymenoptera
Hymenoptera is a large order (biology), order of insects, comprising the sawfly, sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are Parasitoid wasp, parasitic. Females typically have a special ovipositor for inserting eggs into hosts or places that are otherwise inaccessible. This ovipositor is often modified into a stinger. The young develop through holometabolism (complete metamorphosis (biology), metamorphosis)—that is, they have a wormlike larval stage and an inactive pupal stage before they mature. Etymology The name Hymenoptera refers to the wings of the insects, but the original derivation is ambiguous. All references agree that the derivation involves the Ancient Greek language, Ancient Greek wikt:πτερόν, πτερόν (''pteron'') for wing. The Ancient Greek wikt:ὑμήν, ὑμήν (''hymen'') for membrane provides a plausible etymology for the term bec ...
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Xyeloidea
The Xyelidae are a comparatively species-poor family of sawflies comprising about 80 extant species in five genera worldwide and is the only family in the superfamily Xyeloidea. The fossil record of the family is extensive, comprising more than 120 species and including the oldest fossil Hymenoptera species dating back to the Triassic, between 245 and 208 million years ago. Xyelidae are to be regarded as living fossils since they represent one of the oldest lineages of insects and include still extant forms. The extant species occur in the Northern Hemisphere, especially in boreal regions of the Holarctic, though there are a few Oriental species. Two genera and about 15 species occur in Europe.Taeger, A., Blank, S.M. & Liston, A. D. 2006: European Sawflies (Hymenoptera: Symphyta) – A Species Checklist for the Countries. Pp. 399-504. In: Blank, S.M., Schmidt, S. & Taeger, A. (eds): Recent Sawfly Research: Synthesis and Prospects. Goecke & Evers, Keltern.Blank, S.M. 2002: Biosyste ...
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Paraphyly
In taxonomy, a group is paraphyletic if it consists of the group's last common ancestor and most of its descendants, excluding a few monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic group (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of synapomorphies and symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term was coined by Willi Hennig to apply to well-known taxa like Reptilia ( reptiles) which, as commonly named and traditionally defined, is paraphyletic with respect to mammals and birds. Reptilia contains the last common ancestor of reptiles and all descendants of that ancestor, including all extant reptiles as well as the extinct synapsids, ...
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Apocrita
Apocrita is a suborder of insects in the order Hymenoptera. It includes wasps, bees, and ants, and consists of many families. It contains the most advanced hymenopterans and is distinguished from Symphyta by the narrow "waist" ( petiole) formed between the first two segments of the actual abdomen; the first abdominal segment is fused to the thorax, and is called the propodeum. Therefore, it is general practice, when discussing the body of an apocritan in a technical sense, to refer to the mesosoma and metasoma (or gaster) rather than the "thorax" and "abdomen", respectively. The evolution of a constricted waist was an important adaption for the parasitoid lifestyle of the ancestral apocritan, allowing more maneuverability of the female's ovipositor. The ovipositor either extends freely or is retracted, and may be developed into a stinger for both defense and paralyzing prey. Larvae are legless and blind, and either feed inside a host (plant or animal) or in a nest cell provisio ...
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Pamphilioidea
The Pamphilioidea are a small superfamily within the Symphyta (the sawflies), containing some 250 living species restricted to the temperate regions of Eurasia and North America. These hymenopterans share the distinctive feature of a very large, almost prognathous head, which is widest ventrally. The superfamily contains two extant families. The Pamphiliidae are the leaf-rolling or web-spinning sawflies such as ''Acantholyda'', ''Neurotoma'', and ''Pamphilius'' whose larvae eat plants such as conifers; the adults have simple filiform antennae. The Megalodontesidae include genera such as '' Megalodontes'' and several fossil groups. Their larvae eat herbaceous plants, while the adults have serrate or pectinate antennae. References Bibliography * , in Zhang, Z.-Q. (ed.) Animal Biodiversity: An Outline of Higher-level Classification and Survey of Taxonomic Richness (Addenda 2013) * Rasnitsyn, Alexandr P.; Zhang, Haichun & Wang, Bo (2006): Bizarre fossil insects: web-spinning s ...
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Tenthredinoidea
The Tenthredinoidea are the dominant superfamily of sawflies within the Symphyta, containing some 8,400 species worldwide, primarily in the family Tenthredinidae. All known larvae are phytophagous, and a number are considered pests. The included extant families share the distinctive features of a medially narrowed pronotum, paired protibial spurs, and the loss of the transverse mesonotal groove. The superfamily also includes two extinct families. Meicai and Haiyan (1998) identified 66 extant tribes and 17 subfamilies. Taxonomy Families *Argidae Konow, 1890 (58 genera, 897 spp.) * Blasticotomidae Thomson, 1871 (3 genera, 13 spp.) * Cimbicidae W. Kirby, 1837 (16 genera, 182 spp.) *Diprionidae Rohwer, 1910 (11 genera, 136 spp.) * Pergidae Rohwer, 1911 (60 genera, 442 spp.) *Tenthredinidae Tenthredinidae is the largest family of sawflies, with well over 7,500 species worldwide, divided into 430 genera. Larvae are herbivores and typically feed on the foliage of trees a ...
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Cephoidea
Cephoidea is a small superfamily within the Symphyta, commonly referred to as stem sawflies, containing some 100 species in 10 genera in the living family, Cephidae, plus another 17 genera in the extinct family Sepulcidae. Most species occur in the Northern Hemisphere, especially in Eurasia. The larvae are stem borers in various plants, especially grasses, but sometimes other herbaceous plants, shrubs, or trees. A few are pests of cereal grains (e.g. ''Cephus cinctus'', which attacks wheat). They are exceptionally slender for symphytans, often resembling other types of wasps, and they are the only Symphyta which lack cenchri. They are sometimes postulated to be the sister taxon to the Apocrita, though the Orussidae The Orussidae or the parasitic wood wasps represent a small family of sawflies (" Symphyta"). Currently, about 93 extant and four fossil species are known. They take a key position in phylogenetic analyses of Hymenoptera, because they form the s ... are more common ...
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Karatavitoidea
Karatavitidae is an extinct family of sawflies, known from the Jurassic period, they are the only members of the superfamily Karatavitoidea. While once proposed to be grouped with the Orussoidea in the infraorder Orussomorpha, they are now considered to be the closest relatives of clade containing Orussoidea and Apocrita. There are about 7 genera in Karatavitidae. Genera These seven genera belong to the family Karatavitidae: * † '' Grimmaratavites'' Rasnitsyn, Ansorge & Zhang, 2006 Green Series, Germany, Early Jurassic (Toarcian) * † '' Karatavites'' Rasnitsyn, 1963 Karabastau Formation Kazakhstan, Middle-Late Jurassic (Callovian/ Oxfordian), Daohugou The Haifanggou Formation (), also known as the Jiulongshan Formation (), is a fossil-bearing rock deposit located near Daohugou () village of Ningcheng County, in Inner Mongolia, northeastern China. The formation consists of coarse conglomerates ..., China, Callovian * † '' Postxiphydria'' Rasnitsyn & Zhang, 2010 Karabast ...
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Ovipositor
The ovipositor is a tube-like organ used by some animals, especially insects, for the laying of eggs. In insects, an ovipositor consists of a maximum of three pairs of appendages. The details and morphology of the ovipositor vary, but typically its form is adapted to functions such as preparing a place for the egg, transmitting the egg, and then placing it properly. For most insects, the organ is used merely to attach the egg to some surface, but for many parasitic species (primarily in wasps and other Hymenoptera), it is a piercing organ as well. Some ovipositors only retract partly when not in use, and the basal part that sticks out is known as the scape, or more specifically oviscape, the word ''scape'' deriving from the Latin word '' scāpus'', meaning "stalk" or "shaft". In insects Grasshoppers use their ovipositors to force a burrow into the earth to receive the eggs. Cicadas pierce the wood of twigs with their ovipositors to insert the eggs. Sawflies slit the ...
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Abdomen
The abdomen (colloquially called the belly, tummy, midriff, tucky or stomach) is the part of the body between the thorax (chest) and pelvis, in humans and in other vertebrates. The abdomen is the front part of the abdominal segment of the torso. The area occupied by the abdomen is called the abdominal cavity. In arthropods it is the posterior (anatomy), posterior tagma (biology), tagma of the body; it follows the thorax or cephalothorax. In humans, the abdomen stretches from the thorax at the thoracic diaphragm to the pelvis at the pelvic brim. The pelvic brim stretches from the lumbosacral joint (the intervertebral disc between Lumbar vertebrae, L5 and Vertebra#Sacrum, S1) to the pubic symphysis and is the edge of the pelvic inlet. The space above this inlet and under the thoracic diaphragm is termed the abdominal cavity. The boundary of the abdominal cavity is the abdominal wall in the front and the peritoneal surface at the rear. In vertebrates, the abdomen is a large body c ...
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Parasitoid
In evolutionary ecology, a parasitoid is an organism that lives in close association with its host (biology), host at the host's expense, eventually resulting in the death of the host. Parasitoidism is one of six major evolutionarily stable strategy, evolutionary strategies within parasitism, distinguished by the fatal prognosis for the host, which makes the strategy close to predation. Among parasitoids, strategies range from living inside the host (''endoparasitism''), allowing it to continue growing before emerging as an adult, to Paralysis, paralysing the host and living outside it (''ectoparasitism''). Hosts can include other parasitoids, resulting in hyperparasitism; in the case of oak galls, up to five levels of parasitism are possible. Some parasitoids Behavior-altering parasite, influence their host's behaviour in ways that favour the propagation of the parasitoid. Parasitoids are found in a variety of Taxon, taxa across the insect superorder Endopterygota, whose compl ...
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