Synziphosurine
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Synziphosurine
Synziphosurina is a paraphyletic group of chelicerate arthropods previously thought to be basal horseshoe crabs (Xiphosura). It was later identified as a grade composed of various basal euchelicerates, eventually excluded form the monophyletic Xiphosura ''sensu stricto'' and only regarded as horseshoe crabs under a broader sense ('Xiphosura' ''sensu lato''). Synziphosurines survived at least since early Ordovician to early Carboniferous in ages, with most species are known from the in-between Silurian strata. File:20200920 Synziphosurina dorsal segmentation.png, Dorsal morphology and variation of opisthosomal segmentation across synziphosurines. While ''Weinbergina'' and ''Willwerathia'' being exceptionally large, most synziphosurines are small arthropods with body length ranging only about a few centimeters long. The body of synziphosurine composed of a prosoma covered by a dome-like carapace (prosomal dorsal shield) and an opisthosoma with usually unfused 9-11 segments expre ...
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Xiphosura
Xiphosura () is an order of arthropods related to arachnids. They are more commonly known as horseshoe crabs (a name applied more specifically to the only extant family, Horseshoe crab, Limulidae). They first appeared in the Hirnantian (Late Ordovician). Currently, there are only four living species. Xiphosura contains one suborder, Xiphosurida, and several stem-genera. The group has hardly changed in appearance in hundreds of millions of years; the modern horseshoe crabs look almost identical to prehistoric genera and are considered to be living fossils. The most notable difference between ancient and modern forms is that the abdominal segments in present species are fused into a single unit in adults. Xiphosura were historically placed in the class Merostomata, although this term was intended to encompass also the Eurypterida, eurypterids, whence it denoted what is now known to be an unnatural (paraphyletic) group (although this is a grouping recovered in some recent cladistic ...
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Camanchia
''Camanchia'' is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. ''Camanchia'' was regarded as part of the clade Prosomapoda. Fossils of the single and type species, ''C. grovensis'', have been discovered in deposits of the Silurian period in Iowa, in the United States. Alongside '' Venustulus'', ''Camanchia'' is one of the only Silurian synziphosurine with fossil showing evidence of appendages. The prosoma of ''Camanchia'' covered by a smoothly curved subtriangular carapace with broad doublure (ventral thickening run through the margin of carapace). Detail of the 6 prosomal appendage pairs (chelicerae+5 leg pairs) obscure, at least the first leg pair (appendage II) have spur-like terminations. Opisthosoma is externally 10-segmented with tergites possses blunt pleurae (lateral extension). The last 3 opisthosomal segments forming a narrow postabdomen with short pleurae. Telson The telson () is the posterior-most division of the body of an art ...
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Anderella
''Anderella'' is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. ''Anderella'' was regarded as part of the clade Prosomapoda. Fossils of the single and type species, ''A. parva'', have been discovered in deposits of the Carboniferous period in Montana, in the United States. ''Anderella'' is the first and so far (as of 2020) the only Carboniferous synziphosurine being described, making it the youngest member of synziphosurines. ''Anderella'' is also one of the few synziphosurine genera with fossil showing evidence of appendages, but the details are obscure due to their poor preservation. The prosoma of ''Anderella'' possess a suboval carapace slightly longer than the externally 10-segmented opisthosoma (excluding telson). A row of axial nodes run through the opisthosomal tergite A ''tergum'' (Latin for "the back"; plural ''terga'', associated adjective tergal) is the dorsal ('upper') portion of an arthropod segment other than the head. The ante ...
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Willwerathia
''Willwerathia'' is a genus of synziphosurine, a paraphyletic group of horseshoe crab-like fossil chelicerate arthropods. ''Willwerathia'' known only by one species, ''Willwerathia laticeps'', discovered in deposits of the Devonian period from the Klerf Formation, in the Rhenish Slate Mountains of Germany. Morphology As a synziphosurine, ''Willwerathia'' is unusually large and so far the largest known synziphosurine, with largest carapace measured about 90mm in width. Prosoma of ''Willwerathia'' covered by a vaulted carapace with pointed genal spines, recurved (M-shaped) ophthalmic ridges and pairs of dorsal nodes. Tergites of the opisthosoma are either incomplete or disarticulated in available fossil materials, making it difficult to reveal the original number of opisthosomal segments. The opisthosoma of ''Willwerathia'' most likely compose of 10 segments, each expressed by a tergite that bore a median dorsal spine and a pair of tergopleurae (lateral extensions). The opisth ...
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Pasternakevia
''Pasternakevia'' is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. ''Pasternakevia'' was regarded as part of the clade Planaterga. Fossils of the single and type species, ''P. podolica'', have been discovered in deposits of the Silurian period in Podolia, Ukraine. The prosoma of ''Pasternakevia'' covered by a smooth, semicircular carapace with rounded genal cornua (posterolateral corner of carapace). Within the 10-segmented opisthosoma, tergite of the first segment is reduced and usually hidden under the preceding carapace, while the second one is significantly arched and well-developed. All but the first tergite possess well-developed pleurae (lateral extension). A complete telson The telson () is the posterior-most division of the body of an arthropod. Depending on the definition, the telson is either considered to be the final segment of the arthropod body, or an additional division that is not a true segment on accou ... is yet to be ...
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Silurian
The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods (myriapods, arachnids and hexapods) became fully terrestrialized. A significant evolutionary milestone during ...
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Weinbergina Opitzi
''Weinbergina'' is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. Fossils of the single and type species, ''W. opitzi'', have been discovered in deposits of the Devonian period in the HunsrĂĽck Slate, Germany. Morphology ''Weinbergina'' is a relatively large synziphosurine, ranging about 7 cm to 10 cm in full body length. The prosoma is covered by a semicircular carapace with blunt genal cornua (posterolateral corners). There is possible evidence of lateral eyes located just below the ophthalmic ridges. The opisthosoma is externally 10-segmented, expressed by tergites that possess blunt tergopleurae (lateral extension) and axial nodes. However, the opisthosoma is most likely 11-segmented in origin, with the first segment being highly reduced (a synapomorphy of euchelicerates) and possibly covered by the preceding carapace. The last 3 segments form a narrow postabdomen and lacking lateral nodes. Compared to other synziphosurines with ...
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20200929 Synziphosurina Synziphosurine Size Comparison
9 (nine) is the natural number following and preceding . Evolution of the Arabic digit In the beginning, various Indians wrote a digit 9 similar in shape to the modern closing question mark without the bottom dot. The Kshatrapa, Andhra and Gupta started curving the bottom vertical line coming up with a -look-alike. The Nagari continued the bottom stroke to make a circle and enclose the 3-look-alike, in much the same way that the sign @ encircles a lowercase ''a''. As time went on, the enclosing circle became bigger and its line continued beyond the circle downwards, as the 3-look-alike became smaller. Soon, all that was left of the 3-look-alike was a squiggle. The Arabs simply connected that squiggle to the downward stroke at the middle and subsequent European change was purely cosmetic. While the shape of the glyph for the digit 9 has an ascender in most modern typefaces, in typefaces with text figures the character usually has a descender, as, for example, in . The mod ...
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Prosoma
The cephalothorax, also called prosoma in some groups, is a tagma of various arthropods, comprising the head and the thorax fused together, as distinct from the abdomen behind. (The terms ''prosoma'' and ''opisthosoma'' are equivalent to ''cephalothorax'' and ''abdomen'' in some groups.) The word ''cephalothorax'' is derived from the Greek words for head (, ') and thorax (, '). This fusion of the head and thorax is seen in chelicerates and crustaceans; in other groups, such as the Hexapoda (including insects), the head remains free of the thorax. In horseshoe crabs and many crustaceans, a hard shell called the carapace covers the cephalothorax. Arachnid anatomy Fovea The fovea is the centre of the cephalothorax and is located behind the head (only in spiders).Dalton, Steve (2008). ''Spiders; The Ultimate Predators''. A & C Black, London. P.p. 19. . It is often important in identification. It can be transverse or procurved Smith, A. M. (1990c). Baboon spiders: Tarantulas of Afr ...
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Carapace
A carapace is a Dorsum (biology), dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the underside is called the plastron. Crustaceans In crustaceans, the carapace functions as a protective cover over the cephalothorax (i.e., the fused head and thorax, as distinct from the abdomen behind). Where it projects forward beyond the eyes, this projection is called a rostrum (anatomy), rostrum. The carapace is Calcification, calcified to varying degrees in different crustaceans. Zooplankton within the phylum Crustacea also have a carapace. These include Cladocera, ostracods, and Isopoda, isopods, but isopods only have a developed "cephalic shield" carapace covering the head. Arachnids In arachnids, the carapace is formed by the fusion of prosomal tergites into a single Plate (animal anatomy), plate which carries the e ...
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Early Ordovician
The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. The Ordovician, named after the Welsh tribe of the Ordovices, was defined by Charles Lapworth in 1879 to resolve a dispute between followers of Adam Sedgwick and Roderick Murchison, who were placing the same rock beds in North Wales in the Cambrian and Silurian systems, respectively. Lapworth recognized that the fossil fauna in the disputed strata were different from those of either the Cambrian or the Silurian systems, and placed them in a system of their own. The Ordovician received international approval in 1960 (forty years after Lapworth's death), when it was adopted as an official period of the Paleozoic Era by the International Geological Congress. Life continued to flourish during the Ordovician as it did in the earlier Cambr ...
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Opisthosoma
The opisthosoma is the posterior part of the body in some arthropods, behind the prosoma (cephalothorax). It is a distinctive feature of the subphylum Chelicerata (arachnids, horseshoe crabs and others). Although it is similar in most respects to an abdomen (and is often referred to as such), the opisthosoma is differentiated by its inclusion of the respiratory organs (book lungs or book gills) and the heart. Segments The number of segments and appendages on the opisthosoma vary. Scorpions have 13, but the first is only seen during its embryological development. Other arachnids have fewer; harvestmen, for instance, have only ten. In general, appendages are absent or reduced, although in horseshoe crabs they persist as large plate-like limbs, called opercula or branchiophores, bearing the book gills, and that function in locomotion and gas exchange. In most chelicerates the opisthosomal limbs are greatly reduced and persist only as specialized structures, such as the silk-producing ...
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