Camanchia
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Camanchia
''Camanchia'' is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. ''Camanchia'' was regarded as part of the clade Prosomapoda. Fossils of the single and type species, ''C. grovensis'', have been discovered in deposits of the Silurian period in Iowa, in the United States. Alongside '' Venustulus'', ''Camanchia'' is one of the only Silurian synziphosurine with fossil showing evidence of appendages. The prosoma of ''Camanchia'' covered by a smoothly curved subtriangular carapace with broad doublure (ventral thickening run through the margin of carapace). Detail of the 6 prosomal appendage pairs (chelicerae+5 leg pairs) obscure, at least the first leg pair (appendage II) have spur-like terminations. Opisthosoma is externally 10-segmented with tergites possses blunt pleurae (lateral extension). The last 3 opisthosomal segments forming a narrow postabdomen with short pleurae. Telson The telson () is the posterior-most division of the body of an art ...
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Synziphosurine
Synziphosurina is a paraphyletic group of chelicerate arthropods previously thought to be basal horseshoe crabs (Xiphosura). It was later identified as a grade composed of various basal euchelicerates, eventually excluded form the monophyletic Xiphosura ''sensu stricto'' and only regarded as horseshoe crabs under a broader sense ('Xiphosura' ''sensu lato''). Synziphosurines survived at least since early Ordovician to early Carboniferous in ages, with most species are known from the in-between Silurian strata. File:20200920 Synziphosurina dorsal segmentation.png, Dorsal morphology and variation of opisthosomal segmentation across synziphosurines. While ''Weinbergina'' and ''Willwerathia'' being exceptionally large, most synziphosurines are small arthropods with body length ranging only about a few centimeters long. The body of synziphosurine composed of a prosoma covered by a dome-like carapace (prosomal dorsal shield) and an opisthosoma with usually unfused 9-11 segments expre ...
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Synziphosurina
Synziphosurina is a paraphyletic group of chelicerate arthropods previously thought to be basal horseshoe crabs (Xiphosura). It was later identified as a grade composed of various basal euchelicerates, eventually excluded form the monophyletic Xiphosura ''sensu stricto'' and only regarded as horseshoe crabs under a broader sense ('Xiphosura' ''sensu lato''). Synziphosurines survived at least since early Ordovician to early Carboniferous in ages, with most species are known from the in-between Silurian strata. File:20200920 Synziphosurina dorsal segmentation.png, Dorsal morphology and variation of opisthosomal segmentation across synziphosurines. While ''Weinbergina'' and ''Willwerathia'' being exceptionally large, most synziphosurines are small arthropods with body length ranging only about a few centimeters long. The body of synziphosurine composed of a prosoma covered by a dome-like carapace (prosomal dorsal shield) and an opisthosoma with usually unfused 9-11 segments expre ...
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Venustulus
''Venustulus'' is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. ''Venustulus'' was regarded as part of the clade Prosomapoda. Fossils of the single and type species, ''V. waukeshaensis'', have been discovered in deposits of the Silurian period in Wisconsin, in the United States. ''Venustulus'' is one of the few synziphosurine genera with fossil showing evidence of appendages, the other ones being '' Weinbergina, Anderella'' and ''Camanchia''. Despite often being aligned close to Horseshoe crabs, it has been found that ''Venustulus'' and its relatives form a group made up of various basal euchelicerate arthropods more distant to the xiphosurans. Description ''Venustulus'' is the third largest member of its grouping just behind ''Willwerathia'' and ''Weinbergina'' respectively. This arthropod appears to be blind due to the lack of eyes on its carapace, suggesting it lived either in deep water or buried in the sediments. Unlike modern horseshoe ...
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Chelicerate
The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, ticks, and mites, among many others), as well as a number of extinct lineages, such as the eurypterids (sea scorpions) and chasmataspidids. The Chelicerata originated as marine animals in the Middle Cambrian period; the first confirmed chelicerate fossils, belonging to '' Sanctacaris'', date from 508 million years ago. The surviving marine species include the four species of xiphosurans (horseshoe crabs), and possibly the 1,300 species of pycnogonids (sea spiders), if the latter are indeed chelicerates. On the other hand, there are over 77,000 well-identified species of air-breathing chelicerates, and there may be about 500,000 unidentified species. Like all arthropods, chelicerates have segmented bodies with jointed limbs, all covered ...
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Prosomapoda
Prosomapoda is a clade of euchelicerates including the groups Xiphosura (horseshoe crabs) and Planaterga (a group comprising bunodids, pseudoniscids, chasmataspidids, eurypterids and arachnids), as well as several basal synziphosurid genera. The clade is defined by the lack of exopods (outer branches) of prosomal appendage II-V in the adult instar, where in contrast the exopods of appendage II-V are well-developed in the non-prosomapod euchelicerates '' Offacolus'' and ''Dibasterium ''Dibasterium'' is an extinct genus of euchelicerate, a group of chelicerate arthropods. Fossils of the single and type species, ''D. durgae'', have been discovered in the Coalbrookdale Formation of the Middle Silurian period (Homerian age) in ...''. References Middle Ordovician first appearances {{Chelicerata-stub ...
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Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus comes above species and below family (taxonomy), family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants ...
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Silurian Arthropods
The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago (Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods (myriapods, arachnids and hexapods) became fully terrestrialized. A significant evolutionary milestone during the ...
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Silurian First Appearances
The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods (myriapods, arachnids and hexapods) became fully terrestrialized. A significant evolutionary milestone during th ...
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Telson
The telson () is the posterior-most division of the body of an arthropod. Depending on the definition, the telson is either considered to be the final segment of the arthropod body, or an additional division that is not a true segment on account of not arising in the embryo from teloblast areas as other segments. It never carries any appendages, but a forked "tail" called the caudal furca may be present. The shape and composition of the telson differs between arthropod groups. Crustaceans In lobsters, shrimp and other decapods, the telson, along with the uropods, forms the tail fan. This is used as a paddle in the caridoid escape reaction ("lobstering"), whereby an alarmed animal rapidly flexes its tail, causing it to dart backwards. Krill can reach speeds of over 60 cm per second by this means. The trigger time to optical stimulus is, in spite of the low temperatures, only 55 milliseconds. In the Isopoda and Tanaidacea (superorder Peracarida), the last abdominal b ...
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Tergites
A ''tergum'' (Latin for "the back"; plural ''terga'', associated adjective tergal) is the dorsal ('upper') portion of an arthropod segment other than the head. The anterior edge is called the 'base' and posterior edge is called the 'apex' or 'margin'. A given tergum may be divided into hardened plates or sclerites commonly referred to as tergites. In a thoracic segment, for example, the tergum may be divided into an anterior notum and a posterior scutellum. Lateral extensions of a tergite are known as paranota (Greek for "alongside the back") or ''carinae'' (Latin for "keel"), exemplified by the flat-backed millipedes of the order Polydesmida. Kinorhynchs have tergal and sternal plates too, though seemingly not homologous with those of arthropods. Tergo-tergal is a stridulatory mechanism in which fine spines of the abdominal tergites are rubbed together to produce sound. This process is known as abdominal telescoping. Examples File:Andrena spiraeana abdomen.jpg , Abdominal t ...
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Opisthosoma
The opisthosoma is the posterior part of the body in some arthropods, behind the prosoma (cephalothorax). It is a distinctive feature of the subphylum Chelicerata (arachnids, horseshoe crabs and others). Although it is similar in most respects to an abdomen (and is often referred to as such), the opisthosoma is differentiated by its inclusion of the respiratory organs (book lungs or book gills) and the heart. Segments The number of segments and appendages on the opisthosoma vary. Scorpions have 13, but the first is only seen during its embryological development. Other arachnids have fewer; harvestmen, for instance, have only ten. In general, appendages are absent or reduced, although in horseshoe crabs they persist as large plate-like limbs, called opercula or branchiophores, bearing the book gills, and that function in locomotion and gas exchange. In most chelicerates the opisthosomal limbs are greatly reduced and persist only as specialized structures, such as the silk-producing ...
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Chelicerae
The chelicerae () are the mouthparts of the subphylum Chelicerata, an arthropod group that includes arachnids, horseshoe crabs, and sea spiders. Commonly referred to as "jaws", chelicerae may be shaped as either articulated fangs, or similarly to pincers. Some chelicerae, such as those found on nearly all spiders, are hollow and contain (or are connected to) venom glands, and are used to inject venom into prey or a perceived threat. In ''Pisaurina mira'', also known as the nursery web spider, the chelicerae are utilized to snatch the prey once it becomes within reach, facilitating the "sit-and-wait ambush predator" behavior. Both pseudoscorpions and harvestmen have structures on their chelicerae that are used for grooming (papillae in pseudoscorpions, cheliceral teeth in Opiliones). Types Chelicerae can be divided into three kinds: jackknife chelicerae, scissor chelicerae, and 3-segmented chelate chelicerae. Jackknife chelicerae The jackknife chelicera is subchelate (with ...
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