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Sphenodontidae
Sphenodontidae is a family within the reptile group Rhynchocephalia, comprising taxa most closely related to the living tuatara of the genus ''Sphenodon''. Historically the taxa included within Sphenodontidae have varied greatly between analyses, and the group has lacked a formal definition. '' Cynosphenodon'' from the Early Jurassic of Mexico has consistently been recovered as a close relative of the tuatara in most analyses, with the clade containing the two often called Sphenodontinae. The herbivorous Eilenodontinae, otherwise considered part of Opisthodontia, is also sometimes considered part of this family as the sister group to Sphenodontinae. Sphenodontines first appeared during the Early Jurassic, and are characterised by a complete lower temporal bar caused by the fusion of the quadrate/quadratojugal and the jugal The jugal is a skull bone found in most reptiles, amphibians and birds. In mammals, the jugal is often called the malar or zygomatic. It is connected to t ...
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Eilenodontinae
Opisthodontia is a proposed clade of sphenodontian reptiles, uniting ''Opisthias'' from the Late Jurassic-earliest Cretaceous of Europe and North America with the Elienodontinae, a group of herbivorous sphenodontians known from the Late Triassic to Late Cretaceous. Description Teeth and diet Like other sphenodonts, opisthodonts had acrodont teeth which grew directly from the bone. They had one row of teeth on the lower jaw and two rows on the roof of the mouth. When processing food, their mandibular teeth would have slid between the outer (maxillary) teeth and inner (palatine) teeth. Some opisthodonts, such as '' Sphenotitan'', also had clusters of small teeth on the pterygoid at the center of the mouth roof. Opisthodont teeth were wide, numerous, and tightly-packed for grinding and shredding tough plant matter. Although wide shredding teeth are also known in a few other sphenodontians, such as ''Clevosaurus'' and ''Pelecymala'', the most diverse and long-lasting group of h ...
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Rhynchocephalia
Rhynchocephalia (; ) is an order of lizard-like reptiles that includes only one living species, the tuatara (''Sphenodon punctatus'') of New Zealand. Despite its current lack of diversity, during the Mesozoic rhynchocephalians were a diverse group including a wide array of morphologically distinct forms. The oldest record of the group is dated to the Middle Triassic around 238 to 240 million years ago, and they had achieved a worldwide distribution by the Early Jurassic. Most rhynchocephalians belong to the group Sphenodontia ('wedge-teeth'). Their closest living relatives are lizards and snakes in the order Squamata, with the two orders being grouped together in the superorder Lepidosauria. Many of the niches occupied by lizards today were held by sphenodontians during the Triassic and Jurassic, although lizard diversity began to overtake sphenodontian diversity in the Cretaceous, and they had disappeared almost entirely by the beginning of the Cenozoic. While the modern tuat ...
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Opisthodontia (reptile)
Opisthodontia is a proposed clade of sphenodontian reptiles, uniting ''Opisthias'' from the Late Jurassic-earliest Cretaceous of Europe and North America with the Elienodontinae, a group of herbivorous sphenodontians known from the Late Triassic to Late Cretaceous. Description Teeth and diet Like other sphenodonts, opisthodonts had acrodont teeth which grew directly from the bone. They had one row of teeth on the lower jaw and two rows on the roof of the mouth. When processing food, their mandibular teeth would have slid between the outer (maxillary) teeth and inner (palatine) teeth. Some opisthodonts, such as '' Sphenotitan'', also had clusters of small teeth on the pterygoid at the center of the mouth roof. Opisthodont teeth were wide, numerous, and tightly-packed for grinding and shredding tough plant matter. Although wide shredding teeth are also known in a few other sphenodontians, such as ''Clevosaurus'' and ''Pelecymala'', the most diverse and long-lasting group of ...
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Ankylosphenodon
''Ankylosphenodon'' is an extinct species of sphenodontian known from Tepexi de Rodriguez, Mexico. It is known from Early Cretaceous sedimentary deposits from the Tlayua formation. Lifestyle ''Ankylosphenodon'' is thought to have been an aquatic reptile due to its pachyostotic skeleton and other anatomical features often seen in aquatic animals. There is also evidence that its teeth were constantly growing throughout its lifetime, which may be indicative of herbivory, a rare characteristic among Lepidosauria The Lepidosauria (, from Greek meaning ''scaled lizards'') is a subclass or superorder of reptiles, containing the orders Squamata and Rhynchocephalia. Squamata includes snakes, lizards, and amphisbaenians. Squamata contains over 9,000 species .... References Sphenodontia Cretaceous reptiles of North America Prehistoric reptile genera {{Cretaceous-reptile-stub ...
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Cynosphenodon
''Cynosphenodon'' ( ; "Dog Sphenodontian") is an extinct genus of the family Sphenodontidae from the Middle Jurassic La Boca Formation of Tamaulipas, Mexico. Growth patterns in the teeth of ''Cynosphenodon'' suggest its close relationship with the modern tuatara Tuatara (''Sphenodon punctatus'') are reptiles endemic to New Zealand. Despite their close resemblance to lizards, they are part of a distinct lineage, the order Rhynchocephalia. The name ''tuatara'' is derived from the Māori language and m ....Reynoso, V. H. (2003). Growth patterns and ontogenetic variation of the teeth and jaws of the Middle Jurassic sphenodontian ''Cynosphenodon huizachalensis'' (Reptilia: Rhynchocephalia). ''Canadian Journal of Earth Sciences 40''(4), 609–619 References Jurassic lepidosaurs Sphenodontia Fossils of Mexico Prehistoric reptile genera {{Jurassic-reptile-stub ...
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Zapatadon
''Zapatadon'' is an extinct genus of sphenodontid reptile from the end of the Early Jurassic in the lower part of La Boca Formation of Tamaulipas, Mexico.Marisol Montellano, James A. Hopson and James M. Clark (2008)Late Early Jurassic Mammaliaforms from Huizachal Canyon, Tamaulipas, México ''Journal of Vertebrate Paleontology'', Vol. 28, No. 4 (Dec. 12, 2008), pp. 1130-1143. Is known from a nearly complete skull with mandible of a post-hatchling individual (the specimen IGM 3497, in the Instituto de Geologia, of the Universidad Nacional Autónoma de Mexico), and is one of the smallest skulls between the sphenodontians, with an estimated total length of 11.3 millimetres, a bit smaller than the hatchling individuals observed in the modern tuatara (''Sphenodon''); features like the oblique mandibular symphysis suggests that the holotype is from an individual in a relatively mature stage of ontogenic development. ''Zapatadon'' is diagnosed by their hatchling tooth series located in ...
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Kawasphenodon
''Kawasphenodon'' is an extinct genus of sphenodontian reptile, known from the Late Cretaceous and Paleocene of Patagonia in South America. The type species, ''K. expectatus'', was described in 2005 from jaw fragments found in late Campanian aged sediments in the Los Alamitos Formation, the jaw when complete was estimated to be 11 cm long, making it among the largest known sphenodontians. A second species, ''K. peligrensis'', around 1/3 the size of the type species, was described in 2014 also from jaw fragments in early Paleocene (Danian) sediments of the Salamanca Formation, making it the youngest known definitive representative of Rhynchocephalia outside of New Zealand. In the original description, it was found to be a member of Sphenodontidae, in other subsequent analyses it was found to be a member of Opisthodontia. A 2020 analysis of rhyncocephalian relationships found it to be outside Opisthodontia, and instead a member of the Sphenodontinae as the closest known relative ...
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Tuatara
Tuatara (''Sphenodon punctatus'') are reptiles endemic to New Zealand. Despite their close resemblance to lizards, they are part of a distinct lineage, the order Rhynchocephalia. The name ''tuatara'' is derived from the Māori language and means "peaks on the back". The single extant species of tuatara is the only surviving member of its order. Rhynchocephalians originated during the Triassic (~250 million years ago), reached worldwide distribution and peak diversity during the Jurassic and, with the exception of tuatara, were extinct by 60 million years ago. Their closest living relatives are squamates (lizards and snakes). For this reason, tuatara are of interest in the study of the evolution of lizards and snakes, and for the reconstruction of the appearance and habits of the earliest diapsids, a group of amniote tetrapods that also includes dinosaurs (including birds) and crocodilians. Tuatara are greenish brown and grey, and measure up to from head to tail-tip and wei ...
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Tuatara
Tuatara (''Sphenodon punctatus'') are reptiles endemic to New Zealand. Despite their close resemblance to lizards, they are part of a distinct lineage, the order Rhynchocephalia. The name ''tuatara'' is derived from the Māori language and means "peaks on the back". The single extant species of tuatara is the only surviving member of its order. Rhynchocephalians originated during the Triassic (~250 million years ago), reached worldwide distribution and peak diversity during the Jurassic and, with the exception of tuatara, were extinct by 60 million years ago. Their closest living relatives are squamates (lizards and snakes). For this reason, tuatara are of interest in the study of the evolution of lizards and snakes, and for the reconstruction of the appearance and habits of the earliest diapsids, a group of amniote tetrapods that also includes dinosaurs (including birds) and crocodilians. Tuatara are greenish brown and grey, and measure up to from head to tail-tip and wei ...
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Quadrate Bone
The quadrate bone is a skull bone in most tetrapods, including amphibians, sauropsids (reptiles, birds), and early synapsids. In most tetrapods, the quadrate bone connects to the quadratojugal and squamosal bones in the skull, and forms upper part of the jaw joint. The lower jaw articulates at the articular bone, located at the rear end of the lower jaw. The quadrate bone forms the lower jaw articulation in all classes except mammals. Evolutionarily, it is derived from the hindmost part of the primitive cartilaginous upper jaw. Function in reptiles In certain extinct reptiles, the variation and stability of the morphology of the quadrate bone has helped paleontologists in the species-level taxonomy and identification of mosasaur squamates and spinosaurine dinosaurs. In some lizards and dinosaurs, the quadrate is articulated at both ends and movable. In snakes, the quadrate bone has become elongated and very mobile, and contributes greatly to their ability to swallow very ...
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Early Jurassic
The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-Jurassic extinction event, 201.3 Ma (million years ago), and ends at the start of the Middle Jurassic 174.1 Ma. Certain rocks of marine origin of this age in Europe are called "Lias Group, Lias" and that name was used for the period, as well, in 19th-century geology. In southern Germany rocks of this age are called Black Jurassic. Origin of the name Lias There are two possible origins for the name Lias: the first reason is it was taken by a geologist from an England, English quarryman's dialect pronunciation of the word "layers"; secondly, sloops from north Cornwall, Cornish ports such as Bude would sail across the Bristol Channel to the Vale of Glamorgan to load up with rock from coastal limestone quarries (lias limestone from S ...
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Quadratojugal Bone
The quadratojugal is a skull bone present in many vertebrates, including some living reptiles and amphibians. Anatomy and function In animals with a quadratojugal bone, it is typically found connected to the jugal (cheek) bone from the front and the squamosal bone from above. It is usually positioned at the rear lower corner of the cranium. Many modern tetrapods lack a quadratojugal bone as it has been lost or fused to other bones. Modern examples of tetrapods without a quadratojugal include salamanders, mammals, birds, and squamates (lizards and snakes). In tetrapods with a quadratojugal bone, it often forms a portion of the jaw joint. Developmentally, the quadratojugal bone is a dermal bone in the temporal series, forming the original braincase. The squamosal and quadratojugal bones together form the cheek region and may provide muscular attachments for facial muscles. In reptiles and amphibians In most modern reptiles and amphibians, the quadratojugal is a prominent, strapl ...
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