''Kawasphenodon'' is an extinct genus of

sphenodontian Rhynchocephalia (; ) is an order of lizard-like reptiles that includes only one living species, the tuatara (''Sphenodon punctatus'') of New Zealand. Despite its current lack of diversity, during the Mesozoic rhynchocephalians were a diverse g ...

reptile, known from the

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...

and

Paleocene The Paleocene, ( ) or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), E ...

of Patagonia in South America. The type species, ''K. expectatus'', was described in 2005 from jaw fragments found in late Campanian aged sediments in the Los Alamitos Formation, the jaw when complete was estimated to be 11 cm long, making it among the largest known sphenodontians. A second species, ''K. peligrensis'', around 1/3 the size of the type species, was described in 2014 also from jaw fragments in early

(

Danian The Danian is the oldest age or lowest stage of the Paleocene Epoch or Series, of the Paleogene Period or System, and of the Cenozoic Era or Erathem. The beginning of the Danian (and the end of the preceding Maastrichtian) is at the Cretaceous� ...

) sediments of the Salamanca Formation, making it the youngest known definitive representative of Rhynchocephalia outside of New Zealand. In the original description, it was found to be a member of

Sphenodontidae Sphenodontidae is a family within the reptile group Rhynchocephalia, comprising taxa most closely related to the living tuatara of the genus ''Sphenodon''. Historically the taxa included within Sphenodontidae have varied greatly between analyses ...

, in other subsequent analyses it was found to be a member of Opisthodontia. A 2020 analysis of rhyncocephalian relationships found it to be outside Opisthodontia, and instead a member of the Sphenodontinae as the closest known relative of the

tuatara Tuatara (''Sphenodon punctatus'') are reptiles endemic to New Zealand. Despite their close resemblance to lizards, they are part of a distinct lineage, the order Rhynchocephalia. The name ''tuatara'' is derived from the Māori language and m ...

, with an estimated divergence between the two genera in the Early Cretaceous. Like most other rhynchocephalians, the teeth are acrodont, with a deep dentary, and it probably had an omnivorous habit.

References

{{Taxonbar, from=Q11737548 Sphenodontia Cretaceous reptiles of South America Paleocene reptiles of South America Cretaceous Argentina Paleogene Argentina Fossils of Argentina Tiupampan Peligran Campanian first appearances Paleocene extinctions Fossil taxa described in 2005 Golfo San Jorge Basin Salamanca Formation