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Spermosphere
In plant science, the spermosphere is the zone in the soil surrounding a germinating seed. This is a small volume with radius perhaps 1 cm but varying with seed type, the variety of soil microorganisms, the level of soil moisture, and other factors. Within the spermosphere a range of complex interactions take place among the germinating seed, the soil, and the microbiome. Because germination is a brief process, the spermosphere is transient, but the impact of the microbial activity within the spermosphere can have strong and long-lasting effects on the developing plant. Seeds exude various molecules that influence their surrounding microbial communities, either inhibiting or stimulating their growth. The composition of the exudates varies according to the plant type and such properties of the soil as its pH and moisture content. With these biochemical effects, the spermosphere develops both downward—to form the rhizosphere (upon the emergence of the plant's radicle)—and u ...
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Exudate
An exudate is a fluid emitted by an organism through pores or a wound, a process known as exuding or exudation. ''Exudate'' is derived from ''exude'' 'to ooze' from Latin ''exsūdāre'' 'to (ooze out) sweat' (''ex-'' 'out' and ''sūdāre'' 'to sweat'). Medicine An exudate is any fluid that filters from the circulatory system into lesions or areas of inflammation. It can be a pus-like or clear fluid. When an injury occurs, leaving skin exposed, it leaks out of the blood vessels and into nearby tissues. The fluid is composed of serum, fibrin, and leukocytes. Exudate may ooze from cuts or from areas of infection or inflammation. Types * Purulent or suppurative exudate consists of plasma with both active and dead neutrophils, fibrinogen, and necrotic parenchymal cells. This kind of exudate is consistent with more severe infections, and is commonly referred to as pus. * Fibrinous exudate is composed mainly of fibrinogen and fibrin. It is characteristic of rheumatic carditis, ...
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Laimosphere
The laimosphere is the microbiologically enriched zone of soil that surrounds below-ground portions of plant stems; the laimosphere is analogous to the rhizosphere and spermosphere. The combining form laim- from (Greek: λαιμός, "throat") denotes a connecting organ (neck) while -sphere indicates a zone of influence. Topographically, the ''laimosphere'' includes the soil around any portion of subterranean plant organs other than roots where exuded nutrients (especially sugars and amino acids) stimulate microbial activities. Subterranean plant organs with a laimosphere include hypocotyls, epicotyls, stems, stolons, corms, bulbs, and leaves. Propagules of soil-borne plant pathogens, whose germination is stimulated by a plant exudates in the laimosphere, can initiate hypocotyl and stem rots leading to "damping-off". Pathogens commonly found to cause such diseases are species of ''Fusarium'', ''Phoma'', ''Phytophthora'', ''Pythium'', ''Rhizoctonia'' and ''Sclerotinia ''Scle ...
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Radicle
In botany, the radicle is the first part of a seedling (a growing plant embryo) to emerge from the seed during the process of germination. The radicle is the embryonic root of the plant, and grows downward in the soil (the shoot emerges from the plumule). Above the radicle is the embryonic stem or hypocotyl, supporting the cotyledon(s). It is the embryonic root inside the seed. It is the first thing to emerge from a seed and down into the ground to allow the seed to suck up water and send out its leaves so that it can start photosynthesizing. The radicle emerges from a seed through the micropyle. Radicles in seedlings are classified into two main types. Those pointing away from the seed coat scar or hilum are classified as ''antitropous'', and those pointing towards the hilum are ''syntropous''. If the radicle begins to decay, the seedling undergoes pre-emergence damping off. This disease appears on the radicle as darkened spots. Eventually, it causes death of the se ...
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Plant Science
Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek word (''botanē'') meaning " pasture", " herbs" " grass", or " fodder"; is in turn derived from (), "to feed" or "to graze". Traditionally, botany has also included the study of fungi and algae by mycologists and phycologists respectively, with the study of these three groups of organisms remaining within the sphere of interest of the International Botanical Congress. Nowadays, botanists (in the strict sense) study approximately 410,000 species of land plants of which some 391,000 species are vascular plants (including approximately 369,000 species of flowering plants), and approximately 20,000 are bryophytes. Botany originated in prehistory as herbalism with the efforts of early humans to identify – and later cultivate – edible, ...
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Germination
Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spores of fungi, ferns, bacteria, and the growth of the pollen tube from the pollen grain of a seed plant. Seed plants Germination is usually the growth of a plant contained within a seed; it results in the formation of the seedling. It is also the process of reactivation of metabolic machinery of the seed resulting in the emergence of radicle and plumule. The seed of a vascular plant is a small package produced in a fruit or cone after the union of male and female reproductive cells. All fully developed seeds contain an embryo and, in most plant species some store of food reserves, wrapped in a seed coat. Some plants produce varying numbers of seeds that lack embryos; these are empty seeds which never germinate. Dormant seeds are viable seeds th ...
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Microorganism
A microorganism, or microbe,, ''mikros'', "small") and ''organism'' from the el, ὀργανισμός, ''organismós'', "organism"). It is usually written as a single word but is sometimes hyphenated (''micro-organism''), especially in older texts. The informal synonym ''microbe'' () comes from μικρός, mikrós, "small" and βίος, bíos, "life". is an organism of microscopic size, which may exist in its single-celled form or as a colony of cells. The possible existence of unseen microbial life was suspected from ancient times, such as in Jain scriptures from sixth century BC India. The scientific study of microorganisms began with their observation under the microscope in the 1670s by Anton van Leeuwenhoek. In the 1850s, Louis Pasteur found that microorganisms caused food spoilage, debunking the theory of spontaneous generation. In the 1880s, Robert Koch discovered that microorganisms caused the diseases tuberculosis, cholera, diphtheria, and anthrax. Because ...
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Microbiome
A microbiome () is the community of microorganisms that can usually be found living together in any given habitat. It was defined more precisely in 1988 by Whipps ''et al.'' as "a characteristic microbial community occupying a reasonably well-defined habitat which has distinct physio-chemical properties. The term thus not only refers to the microorganisms involved but also encompasses their theatre of activity". In 2020, an international panel of experts published the outcome of their discussions on the definition of the microbiome. They proposed a definition of the microbiome based on a revival of the "compact, clear, and comprehensive description of the term" as originally provided by Whipps ''et al.'', but supplemented with two explanatory paragraphs. The first explanatory paragraph pronounces the dynamic character of the microbiome, and the second explanatory paragraph clearly separates the term ''microbiota'' from the term ''microbiome''. The microbiota consists of al ...
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Rhizosphere
The rhizosphere is the narrow region of soil or substrate that is directly influenced by root secretions and associated soil microorganisms known as the root microbiome. Soil pores in the rhizosphere can contain many bacteria and other microorganisms that feed on sloughed-off plant cells, termed ''rhizodeposition'', and the proteins and sugars released by roots, termed root exudates. This symbiosis leads to more complex interactions, influencing plant growth and competition for resources. Much of the nutrient cycling and disease suppression by antibiotics required by plants, occurs immediately adjacent to roots due to root exudates and metabolic products of symbiotic and pathogenic communities of microorganisms. The rhizosphere also provides space to produce allelochemicals to control neighbours and relatives. The ''rhizoplane'' refers to the root surface including its associated soil particles which closely interact with each other. The plant-soil feedback loop and other phy ...
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Plant Stem
A stem is one of two main structural axes of a vascular plant, the other being the root. It supports leaves, flowers and fruits, transports water and dissolved substances between the roots and the shoots in the xylem and phloem, stores nutrients, and produces new living tissue. The stem can also be called halm or haulm. The stem is normally divided into nodes and internodes: * The nodes hold one or more leaves, as well as buds which can grow into branches (with leaves, conifer cones, or flowers). Adventitious roots may also be produced from the nodes. * The internodes distance one node from another. The term " shoots" is often confused with "stems"; "shoots" generally refers to new fresh plant growth including both stems and other structures like leaves or flowers. In most plants stems are located above the soil surface but some plants have underground stems. Stems have four main functions which are: * Support for and the elevation of leaves, flowers, and fruits. The ...
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Soil Biology
Soil biology is the study of microbial and faunal activity and ecology in soil. Soil life, soil biota, soil fauna, or edaphon is a collective term that encompasses all organisms that spend a significant portion of their life cycle within a soil profile, or at the soil-litter interface. These organisms include earthworms, nematodes, protozoa, fungi, bacteria, different arthropods, as well as some reptiles (such as snakes), and species of burrowing mammals like gophers, moles and prairie dogs. Soil biology plays a vital role in determining many soil characteristics. The decomposition of organic matter by soil organisms has an immense influence on soil fertility, plant growth, soil structure, and carbon storage. As a relatively new science, much remains unknown about soil biology and its effect on soil ecosystems. Overview The soil is home to a large proportion of the world's biodiversity. The links between soil organisms and soil functions are complex. The interconnect ...
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Plant Roots
In vascular plants, the roots are the organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often below the surface of the soil, but roots can also be aerial or aerating, that is, growing up above the ground or especially above water. Function The major functions of roots are absorption of water, plant nutrition and anchoring of the plant body to the ground. Anatomy Root morphology is divided into four zones: the root cap, the apical meristem, the elongation zone, and the hair. The root cap of new roots helps the root penetrate the soil. These root caps are sloughed off as the root goes deeper creating a slimy surface that provides lubrication. The apical meristem behind the root cap produces new root cells that elongate. Then, root hairs form that absorb water and mineral nutrients from the soil. The first root in seed producing plants is th ...
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