Prosthecate Bacteria
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Prosthecate Bacteria
Prosthecate bacteria are a non-phylogenetically related group of Gram-negative bacteria that possess appendages, termed ''prosthecae''. These cellular appendages are neither pili nor flagella, as they are extensions of the cellular membrane and contain cytosol. One notable group of prosthecates is the genus '' Caulobacter''. Function of prostheca Prosthecates are generally chemoorganotrophic aerobes that can grow in nutrient-poor habitats, being able to survive at nutrient levels on the order of parts-per-million for which reason they are often found in aquatic habitats. These bacteria will attach to surfaces with their prosthecae, allowing a greater surface area with which to take up nutrients (and release waste products). Some prosthecates will grow in nutrient-poor soils as aerobic heterotrophs. See also '' Caulobacter'' Oligotrophic Flagella Pilus A pilus (Latin for 'hair'; plural: ''pili'') is a hair-like appendage found on the surface of many bacteria and archaea. ...
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Phylogenetically
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does n ...
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Gram-negative
Gram-negative bacteria are bacteria that do not retain the crystal violet stain used in the Gram staining method of bacterial differentiation. They are characterized by their cell envelopes, which are composed of a thin peptidoglycan cell wall sandwiched between an inner cytoplasmic cell membrane and a bacterial outer membrane. Gram-negative bacteria are found in virtually all environments on Earth that support life. The gram-negative bacteria include the model organism ''Escherichia coli'', as well as many pathogenic bacteria, such as ''Pseudomonas aeruginosa'', ''Chlamydia trachomatis'', and ''Yersinia pestis''. They are a significant medical challenge as their outer membrane protects them from many antibiotics (including penicillin), detergents that would normally damage the inner cell membrane, and lysozyme, an antimicrobial enzyme produced by animals that forms part of the innate immune system. Additionally, the outer leaflet of this membrane comprises a complex lipo ...
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Appendages
An appendage (or outgrowth) is an external body part, or natural prolongation, that protrudes from an organism's body. In arthropods, an appendage refers to any of the homologous body parts that may extend from a body segment, including antennae, mouthparts (including mandibles, maxillae and maxillipeds), gills, locomotor legs (pereiopods for walking, and pleopods for swimming), sexual organs (gonopods), and parts of the tail (uropods). Typically, each body segment carries one pair of appendages. An appendage which is modified to assist in feeding is known as a maxilliped or gnathopod. In vertebrates, an appendage can refer to a locomotor part such as a tail, fins on a fish, limbs (legs, flippers or wings) on a tetrapod; exposed sex organ; defensive parts such as horns and antlers; or sensory organs such as auricles, proboscis ( trunk and snout) and barbels. Appendages may become ''uniramous'', as in insects and centipedes, where each appendage comprises a single series ...
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Pilus
A pilus (Latin for 'hair'; plural: ''pili'') is a hair-like appendage found on the surface of many bacteria and archaea. The terms ''pilus'' and '' fimbria'' (Latin for 'fringe'; plural: ''fimbriae'') can be used interchangeably, although some researchers reserve the term ''pilus'' for the appendage required for bacterial conjugation. All conjugative pili are primarily composed of pilin – fibrous proteins, which are oligomeric. Dozens of these structures can exist on the bacterial and archaeal surface. Some bacteria, viruses or bacteriophages attach to receptors on pili at the start of their reproductive cycle. Pili are antigenic. They are also fragile and constantly replaced, sometimes with pili of different composition, resulting in altered antigenicity. Specific host responses to old pili structures are not effective on the new structure. Recombination genes of pili code for variable (V) and constant (C) regions of the pili (similar to immunoglobulin diversity). As the ...
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Flagella
A flagellum (; ) is a hairlike appendage that protrudes from certain plant and animal sperm cells, and from a wide range of microorganisms to provide motility. Many protists with flagella are termed as flagellates. A microorganism may have from one to many flagella. A gram-negative bacterium ''Helicobacter pylori'' for example uses its multiple flagella to propel itself through the mucus lining to reach the stomach epithelium, where it may cause a gastric ulcer to develop. In some bacteria the flagellum can also function as a sensory organelle, being sensitive to wetness outside the cell. Across the three domains of Bacteria, Archaea, and Eukaryota the flagellum has a different structure, protein composition, and mechanism of propulsion but shares the same function of providing motility. The Latin word means " whip" to describe its lash-like swimming motion. The flagellum in archaea is called the archaellum to note its difference from the bacterial flagellum. Eukaryoti ...
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Cellular Membrane
The cell membrane (also known as the plasma membrane (PM) or cytoplasmic membrane, and historically referred to as the plasmalemma) is a biological membrane that separates and protects the interior of all cells from the outside environment (the extracellular space). The cell membrane consists of a lipid bilayer, made up of two layers of phospholipids with cholesterols (a lipid component) interspersed between them, maintaining appropriate membrane fluidity at various temperatures. The membrane also contains membrane proteins, including integral proteins that span the membrane and serve as membrane transporters, and peripheral proteins that loosely attach to the outer (peripheral) side of the cell membrane, acting as enzymes to facilitate interaction with the cell's environment. Glycolipids embedded in the outer lipid layer serve a similar purpose. The cell membrane controls the movement of substances in and out of cells and organelles, being selectively permeable to ions an ...
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Cytosol
The cytosol, also known as cytoplasmic matrix or groundplasm, is one of the liquids found inside cells (intracellular fluid (ICF)). It is separated into compartments by membranes. For example, the mitochondrial matrix separates the mitochondrion into many compartments. In the eukaryotic cell, the cytosol is surrounded by the cell membrane and is part of the cytoplasm, which also comprises the mitochondria, plastids, and other organelles (but not their internal fluids and structures); the cell nucleus is separate. The cytosol is thus a liquid matrix around the organelles. In prokaryotes, most of the chemical reactions of metabolism take place in the cytosol, while a few take place in membranes or in the periplasmic space. In eukaryotes, while many metabolic pathways still occur in the cytosol, others take place within organelles. The cytosol is a complex mixture of substances dissolved in water. Although water forms the large majority of the cytosol, its structure and prope ...
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Caulobacter
''Caulobacter'' is a genus of Gram-negative bacteria in the class Alphaproteobacteria. Its best-known member is ''Caulobacter crescentus'', an organism ubiquitous in freshwater lakes and rivers; many members of the genus are specialized to Trophic state index#Oligotrophic, oligotrophic environments. Interactions with other organisms Pathogenicity Although ''Caulobacter'' is not commonly appreciated as a cause of human diseases, ''Caulobacter'' isolates have been implicated in a number of cases of recurrent peritonitis in peritoneal dialysis patients. One study has identified the species ''C. crescentus'' and ''C. mirare'' as the cause of a disease of the moth ''Galleria mellonella''; the absence of identified distinct virulence factors in ''C. mirare'' may suggest that other ''Caulobacter'' species have pathogenic potential. References External links

* https://microbewiki.kenyon.edu/index.php/Caulobacter * https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Undef& ...
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Chemoorganotroph
Primary nutritional groups are groups of organisms, divided in relation to the nutrition mode according to the sources of energy and carbon, needed for living, growth and reproduction. The sources of energy can be light or chemical compounds; the sources of carbon can be of organic or inorganic origin. The terms ''aerobic respiration'', ''anaerobic respiration'' and ''fermentation'' (''substrate-level phosphorylation'') do not refer to primary nutritional groups, but simply reflect the different use of possible electron acceptors in particular organisms, such as O2 in aerobic respiration, or nitrate (), sulfate () or fumarate in anaerobic respiration, or various metabolic intermediates in fermentation. Primary sources of energy ''Phototrophs'' absorb light in photoreceptors and transform it into chemical energy. ''Chemotrophs'' release chemical energy. The freed energy is stored as potential energy in ATP, carbohydrates, or proteins. Eventually, the energy is used for life ...
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Heterotrophs
A heterotroph (; ) is an organism that cannot produce its own food, instead taking nutrition from other sources of organic carbon, mainly plant or animal matter. In the food chain, heterotrophs are primary, secondary and tertiary consumers, but not producers. Living organisms that are heterotrophic include all animals and fungi, some bacteria and protists, and many parasitic plants. The term heterotroph arose in microbiology in 1946 as part of a classification of microorganisms based on their type of nutrition. The term is now used in many fields, such as ecology in describing the food chain. Heterotrophs may be subdivided according to their energy source. If the heterotroph uses chemical energy, it is a chemoheterotroph (e.g., humans and mushrooms). If it uses light for energy, then it is a photoheterotroph (e.g., green non-sulfur bacteria). Heterotrophs represent one of the two mechanisms of nutrition (trophic levels), the other being autotrophs (''auto'' = self, ''troph'' ...
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Oligotroph
An oligotroph is an organism that can live in an environment that offers very low levels of nutrients. They may be contrasted with copiotrophs, which prefer nutritionally rich environments. Oligotrophs are characterized by slow growth, low rates of metabolism, and generally low population density. Oligotrophic environments are those that offer little to sustain life. These environments include deep oceanic sediments, caves, glacial and polar ice, deep subsurface soil, aquifers, ocean waters, and leached soils. Examples of oligotrophic organisms are the cave-dwelling olm; the bacterium " ''Candidatus'' Pelagibacter communis", which is the most abundant organism in the oceans with an estimated 2 × 1028 individuals in total; and the lichens with their extremely low metabolic rate. Etymologically, the word "oligotroph" is a combination of the Greek adjective ''oligos'' (ὀλίγος) meaning "few" and the adjective ''trophikos'' (τροφικός)) meaning "feeding". Plant adaptatio ...
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