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Proto-mitochondrion
The proto-mitochondrion is the hypothetical ancestral bacterial endosymbiont from which all mitochondria in eukaryotes are thought to descend, after an episode of symbiogenesis which created the aerobic eukaryotes. Phylogeny The phylogenetic analyses of the few genes that are still encoded in the genomes of modern mitochondria suggest an alphaproteobacterial origin for this endosymbiont, in an ancient episode of symbiogenesis early in the history of the eukaryotes. Although the order '' Rickettsiales'' has been proposed as the alphaproteobacterial sister-group of mitochondria, there is no definitive evidence as to which alphaproteobacterial group the proto-mitochondrion emerged from. Martijn ''et al'' found mitochondria are a possible sister-group to all other alphaproteobacteria. The phylogenetic tree of the Rickettsidae has been inferred by Ferla ''et al.'' from the comparison of 16S + 23S ribosomal RNA sequences. Metabolism Toni Gabaldón and Martijn Huynen (2003) r ...
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Symbiogenesis
Symbiogenesis (endosymbiotic theory, or serial endosymbiotic theory,) is the leading evolutionary theory of the origin of eukaryotic cells from prokaryotic organisms. The theory holds that mitochondria, plastids such as chloroplasts, and possibly other organelles of eukaryotic cells are descended from formerly free-living prokaryotes (more closely related to the Bacteria than to the Archaea) taken one inside the other in endosymbiosis. Mitochondria appear to be phylogenetically related to Rickettsiales bacteria, while chloroplasts are thought to be related to cyanobacteria. The idea that chloroplasts were originally independent organisms that merged into a symbiotic relationship with other one-celled organisms dates back to the 19th century, when it was espoused by researchers such as Andreas Schimper. The endosymbiotic theory was articulated in 1905 and 1910 by the Russian botanist Konstantin Mereschkowski, and advanced and substantiated with microbiological evidence ...
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Bacteria
Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were among the first life forms to appear on Earth, and are present in most of its habitats. Bacteria inhabit soil, water, acidic hot springs, radioactive waste, and the deep biosphere of Earth's crust. Bacteria are vital in many stages of the nutrient cycle by recycling nutrients such as the fixation of nitrogen from the atmosphere. The nutrient cycle includes the decomposition of dead bodies; bacteria are responsible for the putrefaction stage in this process. In the biological communities surrounding hydrothermal vents and cold seeps, extremophile bacteria provide the nutrients needed to sustain life by converting dissolved compounds, such as hydrogen sulphide and methane, to energy. Bacteria also live in symbiotic and parasitic re ...
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Sphingomonadales
The ''Sphingomonadales'' are an order of the Alphaproteobacteria. Phylogeny The currently accepted taxonomy is based on the List of Prokaryotic names with Standing in Nomenclature List of Prokaryotic names with Standing in Nomenclature (LPSN) is an online database that maintains information on the naming and taxonomy Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical ... and the phylogeny is based on whole-genome sequences. References Sphingomonadales Bacteria orders Alphaproteobacteria {{Alphaproteobacteria-stub ...
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Midichloriaceae
"''Candidatus'' Midichloriaceae" is a family of bacteria Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were among ..., included in the order Rickettsiales. No member of this family has been cultured in the laboratory, so the entire family has been given the status '' candidatus''. References Rickettsiales {{alphaproteobacteria-stub ...
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Neorickettsia
''Neorickettsia'' is a genus of bacteria. Species or strains in this genus are coccoid or pleomorphic cells that reside in cytoplasmic vacuoles within monocytes and macrophages of dogs, horses, bats, and humans. ''Neorickettsia sennetsu'' causes Sennetsu ehrlichiosis. Unlike other forms of ehrlichiosis, ''Neorickettsia sennetsu'' is transmitted by trematodes from fish. ''Neorickettsia risticii'' causes Potomac horse fever. '' Neorickettsia helminthoeca'' is found in association with the trematode '' Nanophyetus salmincola'', and causes salmon poisoning disease in dogs and other canids. '' Neorickettsia elokominica'' causes a similar disease, Elokomin fluke fever, in canids and other species. Species * ''Neorickettsia risticii'' * ''Neorickettsia sennetsu ''Neorickettsia sennetsu'' is a Gram-negative bacterium that causes Sennetsu ehrlichiosis. See also *''Neorickettsia risticii ''Neorickettsia risticii'', formerly ''Ehrlichia risticii,'' is an obligate intra ...
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Wolbachia
''Wolbachia'' is a genus of intracellular bacteria that infects mainly arthropod species, including a high proportion of insects, and also some nematodes. It is one of the most common parasitic microbes, and is possibly the most common reproductive parasite in the biosphere. Its interactions with its hosts are often complex, and in some cases have evolved to be mutualistic rather than parasitic. Some host species cannot reproduce, or even survive, without ''Wolbachia'' colonisation. One study concluded that more than 16% of neotropical insect species carry bacteria of this genus, and as many as 25 to 70% of all insect species are estimated to be potential hosts. History The genus was first identified in 1924 by Marshall Hertig and Simeon Burt Wolbach in the common house mosquito. They described it as "a somewhat pleomorphic, rodlike, Gram-negative, intracellular organism hatapparently infects only the ovaries and testes". Hertig formally described the species in 19 ...
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Anaplasma
''Anaplasma'' is a genus of bacteria of the alphaproteobacterial order Rickettsiales, family Anaplasmataceae. ''Anaplasma'' species reside in host blood cells and lead to the disease anaplasmosis. The disease most commonly occurs in areas where competent tick vectors are indigenous, including tropical and semitropical areas of the world for intraerythrocytic ''Anaplasma'' spp.Anaplasmas
reviewed and published by , accessed 10 October 2011.
''Anaplasma'' species are biologically transmitted by Ixodes deer-tick vectors, and the prototypical species, ''A. marginale,'' can be mechanically transmitted by biting flies and iatrogenically with blood-contam ...
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Ehrlichia
''Ehrlichia'' is a genus of Rickettsiales bacteria that are transmitted to vertebrates by ticks. These bacteria cause the disease ehrlichiosis, which is considered zoonotic, because the main reservoirs for the disease are animals. ''Ehrlichia'' species are obligately intracellular pathogens and are transported between cells through the host cell filopodia during initial stages of infection, whereas in the final stages of infection, the pathogen ruptures the host cell membrane. History The genus ''Ehrlichia'' is named after German microbiologist Paul Ehrlich. The first ehrlichial disease was recognized in South Africa during the 19th century. Its tick-borne nature was determined in 1900. The organism itself was demonstrated in 1925 when it was recognized to be a ''Rickettsia''. It was initially named ''Rickettsia ruminantium'', and is currently named '' Ehrlichia ruminantium''. In 1945, an "infection and treatment" method for livestock was developed. This is still the only com ...
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Anaplasmataceae
The Ehrlichiaceae are a family of bacteria, included in the order Rickettsiales The Rickettsiales, informally called rickettsias, are an order of small Alphaproteobacteria. They are obligate intracellular parasites, and some are notable pathogens, including '' Rickettsia'', which causes a variety of diseases in humans, and .... References External links * Rickettsiales Bacteria families {{alphaproteobacteria-stub ...
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Pelagibacter
"''Candidatus'' Pelagibacter", with the single species "''Ca.'' P. communis", was isolated in 2002 and given a specific name, although it has not yet been described as required by the bacteriological code. It is an abundant member of the SAR11 clade in the phylum ''Alphaproteobacteria''. SAR11 members are highly dominant organisms found in both salt and fresh water worldwide and were originally known only from their rRNA genes, first identified in the Sargasso Sea in 1990 by Stephen Giovannoni's laboratory at Oregon State University and later found in oceans worldwide. "''Ca.'' P. communis" and its relatives may be the most abundant organisms in the ocean, and quite possibly the most abundant bacteria in the entire world. It can make up about 25% of all microbial plankton cells, and in the summer they may account for approximately half the cells present in temperate ocean surface water. The total abundance of "''Ca.'' P. communis" and relatives is estimated to be about 2 ×& ...
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Pelagibacteraceae
The Pelagibacteraceae are a family in the Alphaproteobacteria composed of free-living marine bacteria Marine prokaryotes are marine bacteria and marine archaea. They are defined by their habitat as prokaryotes that live in marine environments, that is, in the saltwater of seas or oceans or the brackish water of coastal estuaries. All cellula .... References Alphaproteobacteria {{alphaproteobacteria-stub ...
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Pelagibacterales
The Pelagibacterales are an order in the Alphaproteobacteria composed of free-living marine bacteria that make up roughly one in three cells at the ocean's surface. Overall, members of the ''Pelagibacterales'' are estimated to make up between a quarter and a half of all prokaryotic cells in the ocean. Initially, this taxon was known solely by metagenomic data and was known as the SAR11 clade. It was first placed in the Rickettsiales, but was later raised to the rank of order, and then placed as sister order to the Rickettsiales in the subclass Rickettsidae. It includes the highly abundant marine species '' Pelagibacter ubique''. Bacteria in this order are unusually small. Due to their small genome size and limited metabolic function, ''Pelagibacterales'' have become a model organism for 'streamlining theory'. ''P. ubique'' and related species are oligotrophs (scavengers) and feed on dissolved organic carbon and nitrogen. They are unable to fix carbon or nitrogen, but can per ...
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