Male Reproductive Alliances
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Male Reproductive Alliances
Male reproductive alliances can best be understood within the context of traditional male–male competition, as a specific case of cooperative competition. Such cooperative behavior, however, does not necessarily result in the equal sharing of resources among cooperating individuals. Cooperation often requires that individuals decrease their own fitness to increase the fitness of another.Nowak, M. Five rules for the evolution of cooperation. Science 314, 1560–3 (2006).DuVal, E. H. Adaptive advantages of cooperative courtship for subordinate male lance-tailed manakins. ''American Society of Naturalists'' 169, 423–32 (2007). This behavior becomes even more striking when it occurs within the context of cooperative reproduction, where individuals decrease their own reproductive fitness to improve the reproductive fitness of another. In some species, males cooperate by forming alliances between related or non-related individuals to gain access to females and prevent other males from ...
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Male–male Competition
Sexual selection is a mode of natural selection in which members of one biological sex choose mates of the other sex to mate with (intersexual selection), and compete with members of the same sex for access to members of the opposite sex (intrasexual selection). These two forms of selection mean that some individuals have greater reproductive success than others within a population, for example because they are more attractive or prefer more attractive partners to produce offspring. Successful males benefit from frequent mating and monopolizing access to one or more fertile females. Females can maximise the return on the energy they invest in reproduction by selecting and mating with the best males. The concept was first articulated by Charles Darwin who wrote of a "second agency" other than natural selection, in which competition between mate candidates could lead to speciation. The theory was given a mathematical basis by Ronald Fisher in the early 20th century. Sexual sele ...
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Inclusive Fitness
In evolutionary biology, inclusive fitness is one of two metrics of evolutionary success as defined by W. D. Hamilton in 1964: * Personal fitness is the number of offspring that an individual begets (regardless of who rescues/rears/supports them) * Inclusive fitness is the number of offspring equivalents that an individual rears, rescues or otherwise supports through its behaviour (regardless of who begets them) An individual's own child, who carries one half of the individual's genes, is defined as one offspring equivalent. A sibling's child, who will carry one-quarter of the individual's genes, is 1/2 offspring equivalent. Similarly, a cousin's child, who has 1/16 of the individual's genes, is 1/8 offspring equivalent. From the gene's point of view, evolutionary success ultimately depends on leaving behind the maximum number of copies of itself in the population. Prior to Hamilton's work, it was generally assumed that genes only achieved this through the number of viable off ...
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Bottlenose Dolphin
Bottlenose dolphins are aquatic mammals in the genus ''Tursiops.'' They are common, cosmopolitan members of the family Delphinidae, the family of oceanic dolphins. Molecular studies show the genus definitively contains two species: the common bottlenose dolphin (''Tursiops truncatus'') and the Indo-Pacific bottlenose dolphin (''Tursiops aduncus''). Others, like the Burrunan dolphin (''Tursiops (aduncus) australis''), may be alternately considered their own species or be subspecies of ''T. aduncus''. Bottlenose dolphins inhabit warm and temperate seas worldwide, being found everywhere except for the Arctic and Antarctic Circle regions. Their name derives from the Latin ''tursio'' (dolphin) and ''truncatus'' for their characteristic truncated teeth. Numerous investigations of bottlenose dolphin intelligence have been conducted, examining mimicry, use of artificial language, object categorization, and self-recognition. They can use tools (sponging; using marine sponges to forage ...
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Lion
The lion (''Panthera leo'') is a large Felidae, cat of the genus ''Panthera'' native to Africa and India. It has a muscular, broad-chested body; short, rounded head; round ears; and a hairy tuft at the end of its tail. It is sexually dimorphic; adult male lions are larger than females and have a prominent mane. It is a social species, forming groups called ''prides''. A lion's pride consists of a few adult males, related females, and cubs. Groups of female lions usually hunt together, preying mostly on large ungulates. The lion is an apex predator, apex and keystone predator; although some lions scavenge when opportunities occur and have been known to hunt Human, humans, lions typically don't actively seek out and prey on humans. The lion inhabits grasslands, savannas and shrublands. It is usually more diurnality, diurnal than other wild cats, but when persecuted, it adapts to being active nocturnality, at night and crepuscular, at twilight. During the Neolithic period, the li ...
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Inclusive Fitness
In evolutionary biology, inclusive fitness is one of two metrics of evolutionary success as defined by W. D. Hamilton in 1964: * Personal fitness is the number of offspring that an individual begets (regardless of who rescues/rears/supports them) * Inclusive fitness is the number of offspring equivalents that an individual rears, rescues or otherwise supports through its behaviour (regardless of who begets them) An individual's own child, who carries one half of the individual's genes, is defined as one offspring equivalent. A sibling's child, who will carry one-quarter of the individual's genes, is 1/2 offspring equivalent. Similarly, a cousin's child, who has 1/16 of the individual's genes, is 1/8 offspring equivalent. From the gene's point of view, evolutionary success ultimately depends on leaving behind the maximum number of copies of itself in the population. Prior to Hamilton's work, it was generally assumed that genes only achieved this through the number of viable off ...
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Tit-for-tat
Tit for tat is an English saying meaning "equivalent retaliation". It developed from "tip for tap", first recorded in 1558. It is also a highly effective strategy in game theory. An agent using this strategy will first cooperate, then subsequently replicate an opponent's previous action. If the opponent previously was cooperative, the agent is cooperative. If not, the agent is not. Game theory Tit-for-tat has been very successfully used as a strategy for the iterated prisoner's dilemma. The strategy was first introduced by Anatol Rapoport in Robert Axelrod's two tournaments, held around 1980. Notably, it was (on both occasions) both the simplest strategy and the most successful in direct competition. An agent using this strategy will first cooperate, then subsequently replicate an opponent's previous action. If the opponent previously was cooperative, the agent is cooperative. If not, the agent is not. This is similar to reciprocal altruism in biology. History The term develo ...
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Barbary Macaque
The Barbary macaque (''Macaca sylvanus''), also known as Barbary ape, is a macaque species native to the Atlas Mountains of Algeria, Libya, Tunisia and Morocco, along with a small introduced population in Gibraltar. It is the type species of the genus ''Macaca''. The species is of particular interest because males play an atypical role in rearing young. Because of uncertain paternity, males are integral to raising all infants. Generally, Barbary macaques of all ages and sexes contribute in alloparental care of young. The diet of the Barbary macaque consists primarily of plants and insects and they are found in a variety of habitats. Males live to around 25 years old while females may live up to 30 years. Besides humans, they are the only free-living primates in Europe. Although the species is commonly referred to as the "Barbary ape", the Barbary macaque is actually a true monkey. Its name refers to the Barbary Coast of Northwest Africa. The population of the Barbary macaques i ...
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Lion
The lion (''Panthera leo'') is a large Felidae, cat of the genus ''Panthera'' native to Africa and India. It has a muscular, broad-chested body; short, rounded head; round ears; and a hairy tuft at the end of its tail. It is sexually dimorphic; adult male lions are larger than females and have a prominent mane. It is a social species, forming groups called ''prides''. A lion's pride consists of a few adult males, related females, and cubs. Groups of female lions usually hunt together, preying mostly on large ungulates. The lion is an apex predator, apex and keystone predator; although some lions scavenge when opportunities occur and have been known to hunt Human, humans, lions typically don't actively seek out and prey on humans. The lion inhabits grasslands, savannas and shrublands. It is usually more diurnality, diurnal than other wild cats, but when persecuted, it adapts to being active nocturnality, at night and crepuscular, at twilight. During the Neolithic period, the li ...
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Uganda
}), is a landlocked country in East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the historical .... The country is bordered to the east by Kenya, to the north by South Sudan, to the west by the Democratic Republic of the Congo, to the south-west by Rwanda, and to the south by Tanzania. The southern part of the country includes a substantial portion of Lake Victoria, shared with Kenya and Tanzania. Uganda is in the African Great Lakes region. Uganda also lies within the Nile, Nile basin and has a varied but generally a modified equatorial climate. It has a population of around 49 million, of which 8.5 million live in the Capital city, capital and largest city of Kampala. Uganda is named after the Buganda kingdom, which encompasses a large portion of the south of the country, includi ...
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Kibale National Park
Kibale Forest National Park is a national park in western Uganda, protecting moist evergreen rainforest. It is in size and ranges between and in elevation. Despite encompassing primarily moist evergreen forest, it contains a diverse array of landscapes.McGrew, William, ''et al''. ''Great Ape Societies''. Cambridge University Press, 1996. Print. Kibale is one of the last remaining expanses to contain both lowland and montane forests. In eastern Africa, it sustains the last significant expanse of pre-montane forest. The park was gazetted in 1932 and formally established in 1993 to protect a large area of forest previously managed as a logged forest reserve. The park forms a continuous forest with Queen Elizabeth National Park. This adjoining of the parks creates a wildlife corridor. It is an important ecotourism and safari destination, well-known for its population of habituated chimpanzees and twelve other species of primates. It is also the location of the Makerere University ...
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Savanna Baboon
The yellow baboon (''Papio cynocephalus'') is a baboon in the family of Old World monkeys. The species epithet means "dog-head" in Greek, due to the dog-like shape of the muzzle and head. Yellow baboons have slim bodies with long arms and legs, and yellowish-brown hair. They resemble the Chacma baboon, but are somewhat smaller and with a less elongated muzzle. Their hairless faces are black, framed with white sideburns. Males can grow to about 84 cm, females to about 60 cm. They have long tails which grow to be nearly as long as their bodies. The average life span of the yellow baboon in the wild is roughly 15–20 years; some may live up to 30 years. Yellow baboons inhabit savannas and light forests in eastern Africa, from Kenya and Tanzania to Zimbabwe and Botswana. They are diurnal, terrestrial, and live in complex, mixed-gender social groups of 8 to 200 individuals per troop. Like all other baboon species, they are omnivorous, with a preference for fruits; they also ...
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Cooperation
Cooperation (written as co-operation in British English) is the process of groups of organisms working or acting together for common, mutual, or some underlying benefit, as opposed to working in competition for selfish benefit. Many animal and plant species cooperate both with other members of their own species and with members of other species (symbiosis or mutualism). Among humans Humans cooperate for the same reasons as other animals: immediate benefit, genetic relatedness, and reciprocity, but also for particularly human reasons, such as honesty signaling (indirect reciprocity), cultural group selection, and for reasons having to do with cultural evolution. Language allows humans to cooperate on a very large scale. Certain studies have suggested that fairness affects human cooperation; individuals are willing to punish at their own cost (''altruistic punishment'') if they believe that they are being treated unfairly. Sanfey, et al. conducted an experiment where 19 ind ...
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