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Metriorhynchid
Metriorhynchidae is an extinct family of specialized, aquatic metriorhynchoid crocodyliforms from the Middle Jurassic to the Early Cretaceous period (Bajocian to early Aptian) of Europe, North America and South America. The name Metriorhynchidae was coined by the Austrian zoologist Leopold Fitzinger in 1843.Fitzinger LJFJ. 1843. ''Systema Reptilium''. Wien: Braumüller et Seidel, 106 pp. The group contains two subfamilies, the Metriorhynchinae and the Geosaurinae. They represent the most marine adapted of all archosaurs. Description Metriorhynchids are fully aquatic crocodyliforms. Their forelimbs were small and paddle-like, and unlike living crocodylians, they lost their osteoderms ("armour scutes"). Their body shape maximised hydrodynamy (swimming efficiency), as they did have a shark-like tail fluke. Like ichthyosaurs and plesiosaurs, metriorhynchids developed smooth, scaleless skin. Metriorhynchids were the only group of archosaurs to become fully adapted to the marine real ...
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Cricosaurus
''Cricosaurus'' is an extinct genus of marine crocodyliforms of the Late Jurassic. belonging to the family Metriorhynchidae. The genus was established by Johann Andreas Wagner in 1858 for three skulls from the Tithonian (Late Jurassic) of Germany. The name ''Cricosaurus'' means "Ring lizard", and is derived from the Greek '- ("ring") and -' ("lizard"). It was a relatively small reptile, with ''C. suevicus'' and ''C. araucanensis'' measuring and in total body length, respectively. Discovery ''Cricosaurus'' was first named by Wagner in 1858,Wagner A. 1858. Zur Kenntniss der Saurier aus den lithographischen Schiefern. ''Abhandlungen der Mathemat.-Physikalischen Classe der Königlich Bayerischen Akademie der Wissenschaften'' 8: 415-528. as a reclassification of a specimen he had previously described in 1852 ("Stenosaurus" ''elegans'', "Stenosaurus" being a misspelling of ''Steneosaurus'').Wagner A. 1852. Neu-aufgefundene Saurier-Überreste aus dem lithographischen Schiefern un ...
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Geosaurinae
Geosaurinae is a subfamily of metriorhynchid crocodyliforms from the Middle Jurassic to the Early Cretaceous (Bathonian - Aptian) of Europe, North America and South America. Named by Richard Lydekker, in 1889, it contains the metriorhynchids '' Suchodus'', ''Purranisaurus'', '' Neptunidraco'', '' Tyrannoneustes'', '' Torvoneustes'', ''Dakosaurus'', ''Geosaurus'' and ''Plesiosuchus''. The last four taxa form a tribe within Geosaurinae, the Geosaurini. Geosaurinae is one of two subfamilies of Metriorhynchidae, the other being Metriorhynchinae. These marine reptiles were widespread during the Late Jurassic and Early Cretaceous, their fossilized remains are being frequently found on various places around the world.Daniel Madzia, Sven Sachs, Mark T. Young, Alexander Lukeneder and Petr Skupien (2021)Evidence of two lineages of metriorhynchid crocodylomorphs in the Lower Cretaceous of the Czech Republic ''Acta Palaeontologica Polonica''. doi: https://doi.org/10.4202/app.00801.2020 Phyl ...
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Metriorhynchinae
Metriorhynchinae is a subfamily of metriorhynchid crocodyliforms from the late Middle Jurassic to the Early Cretaceous (Callovian - Valanginian) of Europe, North America and South America. Named by Fitzinger, in 1843, it contains the metriorhynchids '' Maledictosuchus'', ''Gracilineustes'', ''Metriorhynchus'', '' Cricosaurus'' and ''Rhacheosaurus''. The last three taxa form a tribe within Metriorhynchinae, the Rhacheosaurini. Metriorhynchinae is one of two subfamilies of Metriorhynchidae, the other being Geosaurinae. Phylogeny Metriorhynchinae is a stem-based taxon defined in 2009 as the most inclusive clade consisting of ''Metriorhynchus geoffroyii'', but not ''Geosaurus giganteus''. Rhacheosaurini is a stem-based taxon and it was named and defined by Mark T. Young, Mark A. Bell and Stephen L. Brusatte in 2011 as the most inclusive clade including ''Rhacheosaurus gracilis'', but not ''Metriorhynchus geoffroyii'' and ''Gracilineustes leedsi''. The cladogram A cladogram ...
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Archosaur
Archosauria () is a clade of diapsids, with birds and crocodilians as the only living representatives. Archosaurs are broadly classified as reptiles, in the cladistic sense of the term which includes birds. Extinct archosaurs include non-avian dinosaurs, pterosaurs, and extinct relatives of crocodilians. Modern paleontologists define Archosauria as a crown group that includes the most recent common ancestor of living birds and crocodilians, and all of its descendants. The base of Archosauria splits into two clades: Pseudosuchia, which includes crocodilians and their extinct relatives, and Avemetatarsalia, which includes birds and their extinct relatives (such as non-avian dinosaurs and pterosaurs). Older definitions of the group Archosauria rely on shared morphological characteristics, such as an antorbital fenestra in the skull, serrated teeth, and an upright stance. Some extinct reptiles, such as proterosuchids and euparkeriids, possessed these features yet originated pri ...
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Metriorhynchoid
Metriorhynchoidea is an extinct superfamily of thalattosuchian crocodyliforms from the Early Jurassic to the Early Cretaceous (Toarcian - Valanginian, possibly as late as early Aptian) of Europe, North America and South America. Metriorhynchids are fully aquatic crocodyliforms. Named by Fitzinger, in 1843, it contains the basal taxa like ''Teleidosaurus'', ''Zoneait'' and ''Eoneustes'' and the family Metriorhynchidae. An unnamed taxon is known from Chile. Phylogeny Metriorhynchoidea is a stem-based taxon defined in 2009 as the most inclusive clade consisting of ''Metriorhynchus geoffroyii'', but not ''Teleosaurus cadomensis''. The cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ... below follows the topology from a 2011 analysis by Andrea Cau and Federico Fanti. N ...
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Middle Jurassic
The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations containing land animal fossils include the Forest Marble Formation in England, the Kilmaluag Formation in Scotland,British Geological Survey. 2011Stratigraphic framework for the Middle Jurassic strata of Great Britain and the adjoining continental shelf: research report RR/11/06 British Geological Survey, Keyworth, Nottingham. the Daohugou Beds in China, the Itat Formation in Russia, and the Isalo III Formation of western Madagascar. Paleogeography During the Middle Jurassic Epoch, Pangaea began to separate into Laurasia and Gondwana, and the Atlantic Ocean formed. Eastern Laurasia was tectonically active as the Cimmerian plate continued to collide with Laurasia's southern coast, completely closing the Paleo-Tethys Ocean. A subduction zone ...
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Basal (phylogenetics)
In phylogenetics, basal is the direction of the ''base'' (or root) of a phylogenetic tree#Rooted tree, rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, Phylogenetic diversity, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a 'key innovation' implies some degree of correlation between evolutionary innovation and cladogenesis, diversification. However, such a correlation does not make a given ca ...
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Plattenkalk
Plattenkalk is a very finely grained limestone chemically precipitated in a stratified water column under conditions where bioturbation does not occur. The reasons for the quiet depositional environment and the processes of sediment accumulation vary greatly among different plattenkalks, but the lack of bioturbation contributes to the creation of thinly bedded, finely laminated, undisturbed limestones where exceptionally detailed fossils or imprints of organisms occur. The Solnhofen Plattenkalk in Bavaria is a famous example where complete skeletons of large marine vertebrates and impressions of soft-bodied animals have been found. Wilkin, J.T.R. (2020) The south German Plattenkalks. Geology Today 36(1): 27-32. https://doi.org/10.1111/gto.12288Lithographic limestone is a type of plattenkalk chosen for its suitable lithography Lithography () is a planographic method of printing originally based on the immiscibility of oil and water. The printing is from a stone (lithographic ...
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Keichousaurus
''Keichousaurus'' (key-cho-saurus) is a genus of marine reptile in the pachypleurosaur family which went extinct at the close of the Triassic in the Triassic-Jurassic extinction event. The name derives from Kweichow (now Guizhou Province) in China where the first fossil specimen was discovered in 1957. They are among the most common sauropterygian fossils recovered and are often found as nearly complete, articulated skeletons, making them popular among collectors. ''Keichousaurus'', and the pachypleurosaur family broadly, are sometimes classified within Nothosauroidea, but are otherwise listed as a separate, more primitive lineage within Sauropterygia. Description ''Keichousaurus'', like all sauropterygians, was highly adapted to the aquatic environment. Most specimens had small body, males sexually mature with snout-vent length (SVL), and in females by SVL. Mean SVL for mature males is approximately SVL, and for mature females, at most SVL. It had both long necks and lo ...
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Viviparity
Among animals, viviparity is development of the embryo inside the body of the parent. This is opposed to oviparity which is a reproductive mode in which females lay developing eggs that complete their development and hatch externally from the mother. The term 'viviparity' and its adjective form 'viviparous' derive from the Latin ''vivus'' meaning "living" and ''pario'' meaning "give birth to". Reproductive mode Five modes of reproduction have been differentiated in animals based on relations between zygote and parents. The five include two nonviviparous modes: ovuliparity, with external fertilisation, and oviparity, with internal fertilisation. In the latter, the female lays zygotes as eggs with a large vitellus, yolk; this occurs in all birds, most reptiles, and some fishes. These modes are distinguished from viviparity, which covers all the modes that result in live birth: *Histotrophic viviparity: the zygotes develop in the female's oviducts, but find their nutrients b ...
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Histology
Histology, also known as microscopic anatomy or microanatomy, is the branch of biology which studies the microscopic anatomy of biological tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures visible without a microscope. Although one may divide microscopic anatomy into ''organology'', the study of organs, ''histology'', the study of tissues, and ''cytology'', the study of cells, modern usage places all of these topics under the field of histology. In medicine, histopathology is the branch of histology that includes the microscopic identification and study of diseased tissue. In the field of paleontology, the term paleohistology refers to the histology of fossil organisms. Biological tissues Animal tissue classification There are four basic types of animal tissues: muscle tissue, nervous tissue, connective tissue, and epithelial tissue. All animal tissues are considered to be subtypes of these four principal tissue types ...
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