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Geosaurinae
Geosaurinae is a subfamily of metriorhynchid crocodyliforms from the Middle Jurassic to the Early Cretaceous (Bathonian - Aptian) of Europe, North America and South America. Named by Richard Lydekker, in 1889, it contains the metriorhynchids '' Suchodus'', ''Purranisaurus'', '' Neptunidraco'', '' Tyrannoneustes'', '' Torvoneustes'', ''Dakosaurus'', ''Geosaurus'' and ''Plesiosuchus''. The last four taxa form a tribe within Geosaurinae, the Geosaurini. Geosaurinae is one of two subfamilies of Metriorhynchidae, the other being Metriorhynchinae. These marine reptiles were widespread during the Late Jurassic and Early Cretaceous, their fossilized remains are being frequently found on various places around the world.Daniel Madzia, Sven Sachs, Mark T. Young, Alexander Lukeneder and Petr Skupien (2021)Evidence of two lineages of metriorhynchid crocodylomorphs in the Lower Cretaceous of the Czech Republic ''Acta Palaeontologica Polonica''. doi: https://doi.org/10.4202/app.00801.2020 Phyl ...
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Geosaurini
Geosaurinae is a subfamily of metriorhynchid crocodyliforms from the Middle Jurassic to the Early Cretaceous (Bathonian - Aptian) of Europe, North America and South America. Named by Richard Lydekker, in 1889, it contains the metriorhynchids ''Suchodus'', '' Purranisaurus'', '' Neptunidraco'', '' Tyrannoneustes'', '' Torvoneustes'', '' Dakosaurus'', ''Geosaurus'' and '' Plesiosuchus''. The last four taxa form a tribe within Geosaurinae, the Geosaurini. Geosaurinae is one of two subfamilies of Metriorhynchidae, the other being Metriorhynchinae. These marine reptiles were widespread during the Late Jurassic and Early Cretaceous, their fossilized remains are being frequently found on various places around the world.Daniel Madzia, Sven Sachs, Mark T. Young, Alexander Lukeneder and Petr Skupien (2021)Evidence of two lineages of metriorhynchid crocodylomorphs in the Lower Cretaceous of the Czech Republic ''Acta Palaeontologica Polonica''. doi: https://doi.org/10.4202/app.00801.2020 Ph ...
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Metriorhynchid
Metriorhynchidae is an extinct family of specialized, aquatic metriorhynchoid crocodyliforms from the Middle Jurassic to the Early Cretaceous period (Bajocian to early Aptian) of Europe, North America and South America. The name Metriorhynchidae was coined by the Austrian zoologist Leopold Fitzinger in 1843.Fitzinger LJFJ. 1843. ''Systema Reptilium''. Wien: Braumüller et Seidel, 106 pp. The group contains two subfamilies, the Metriorhynchinae and the Geosaurinae. They represent the most marine adapted of all archosaurs. Description Metriorhynchids are fully aquatic crocodyliforms. Their forelimbs were small and paddle-like, and unlike living crocodylians, they lost their osteoderms ("armour scutes"). Their body shape maximised hydrodynamy (swimming efficiency), as they did have a shark-like tail fluke. Like ichthyosaurs and plesiosaurs, metriorhynchids developed smooth, scaleless skin. Metriorhynchids were the only group of archosaurs to become fully adapted to the marine real ...
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Plesiosuchus
''Plesiosuchus'' is an extinct genus of geosaurine metriorhynchid crocodyliform known from the Late Jurassic (late Kimmeridgian to early Tithonian stage) of Dorset, England and possibly also Spain. It contains a single species, ''Plesiosuchus manselii''. Discovery The type and referred specimens of ''Plesiosuchus'' were discovered by John Clavell Mansel-Pleydell in the 1860s alongside the remains of several other large-bodied marine reptiles along the coast of Dorset. Mansel-Pleydell gave these remains to the British Museum (now in the Natural History Museum) in 1866. Part of the holotype of ''P. manselii'' (NHMUK PV OR40103a) was first described by John Hulke in 1869. He referred it to ''Steneosaurus rostro-minor'' Geoffroy (1825), alongside ''Dakosaurus maximus'' and other specimens. Initially, the skull (NHMUK PV OR40103) was believed to be pliosaurian; it was the preparator Mr Davies that suggested a crocodylian nature for the skull. In 1870, Hulke ...
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Dakosaurus Maximus
''Dakosaurus'' is an extinct genus of crocodylomorph within the family Metriorhynchidae that lived during the Late Jurassic and Early Cretaceous. It was large, with teeth that were serrated and compressed lateromedially (flattened from side to side). The genus was established by Friedrich August von Quenstedt in 1856 for an isolated tooth named ''Geosaurus maximus'' by Theodor Plieninger in 1846. ''Dakosaurus'' was a carnivore that spent much, if not all, its life out at sea. The extent of its adaptation to a marine lifestyle means that it is most likely that it mated at sea, but since no eggs or nests have been discovered that have been referred to ''Dakosaurus'', whether it gave birth to live young at sea like dolphins and ichthyosaurs or came ashore like turtles is not known. The name ''Dakosaurus'' means "biter lizard", and is derived from the Greek ' ("biter") and -' ("lizard"). Discovery and species The type species ''Dakosaurus maximus'', meaning "greatest bit ...
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Metriorhynchinae
Metriorhynchinae is a subfamily of metriorhynchid crocodyliforms from the late Middle Jurassic to the Early Cretaceous (Callovian - Valanginian) of Europe, North America and South America. Named by Fitzinger, in 1843, it contains the metriorhynchids '' Maledictosuchus'', ''Gracilineustes'', ''Metriorhynchus'', '' Cricosaurus'' and ''Rhacheosaurus''. The last three taxa form a tribe within Metriorhynchinae, the Rhacheosaurini. Metriorhynchinae is one of two subfamilies of Metriorhynchidae, the other being Geosaurinae. Phylogeny Metriorhynchinae is a stem-based taxon defined in 2009 as the most inclusive clade consisting of ''Metriorhynchus geoffroyii'', but not ''Geosaurus giganteus''. Rhacheosaurini is a stem-based taxon and it was named and defined by Mark T. Young, Mark A. Bell and Stephen L. Brusatte in 2011 as the most inclusive clade including ''Rhacheosaurus gracilis'', but not ''Metriorhynchus geoffroyii'' and ''Gracilineustes leedsi''. The cladogram A cladogram ...
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Dakosaurus
''Dakosaurus'' is an extinct genus of crocodylomorph within the family Metriorhynchidae that lived during the Late Jurassic and Early Cretaceous. It was large, with teeth that were serrated and compressed lateromedially (flattened from side to side). The genus was established by Friedrich August von Quenstedt in 1856 for an isolated tooth named ''Geosaurus maximus'' by Theodor Plieninger in 1846. ''Dakosaurus'' was a carnivore that spent much, if not all, its life out at sea. The extent of its adaptation to a marine lifestyle means that it is most likely that it mated at sea, but since no eggs or nests have been discovered that have been referred to ''Dakosaurus'', whether it gave birth to live young at sea like dolphins and ichthyosaurs or came ashore like turtles is not known. The name ''Dakosaurus'' means "biter lizard", and is derived from the Greek ' ("biter") and -' ("lizard"). Discovery and species The type species ''Dakosaurus maximus'', meaning "greatest biter l ...
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Geosaurus Giganteus
''Geosaurus'' is an extinct genus of marine crocodyliform within the family Metriorhynchidae, that lived during the Late Jurassic and the Early Cretaceous. ''Geosaurus'' was a carnivore that spent much, if not all, its life out at sea. No ''Geosaurus'' eggs or nests have been discovered, so little is known of the reptile's lifecycle, unlike other large marine reptiles of the Mesozoic, such as plesiosaurs or ichthyosaurs which are known to give birth to live young out at sea. Where ''Geosaurus'' mated, whether on land or at sea, is currently unknown. The name ''Geosaurus'' means "Mother of Giants lizard", and is derived from the Greek '' Ge''- ("Earth", the mythical mother of the Giants) and - ("lizard"). The name ''Geosaurus'' was established by the French naturalist Georges Cuvier in 1824.Cuvier G. 1824. Sur les ossements fossiles de crocodiles, 5. In: Dufour & D'Occagne, eds. ''Recherches sur les ossements fossiles, 2nd édition''. Paris: 143-160 Description ''Geosaur ...
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Stem-based Taxon
Phylogenetic nomenclature is a method of nomenclature for taxa in biology that uses phylogenetic definitions for taxon names as explained below. This contrasts with the traditional approach, in which taxon names are defined by a '' type'', which can be a specimen or a taxon of lower rank, and a description in words. Phylogenetic nomenclature is currently regulated by the '' International Code of Phylogenetic Nomenclature'' (''PhyloCode''). Definitions Phylogenetic nomenclature ties names to clades, groups consisting of an ancestor and all its descendants. These groups can equivalently be called monophyletic. There are slightly different ways of specifying the ancestor, which are discussed below. Once the ancestor is specified, the meaning of the name is fixed: the ancestor and all organisms which are its descendants are included in the named taxon. Listing all these organisms (i.e. providing a full circumscription) requires the full phylogenetic tree to be known. In practice, the ...
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Bathonian
In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. Stratigraphic definitions The Bathonian Stage takes its name from Bath, a spa town in England built on Jurassic limestone (the Latinized form of the town name is ''Bathonium''). The name was introduced in scientific literature by Belgian geologist d'Omalius d'Halloy in 1843. The original type locality was located near Bath. The French palaeontologist Alcide d'Orbigny was in 1852 the first to define the exact length of the stage. The base of the Bathonian is at the first appearance of ammonite species '' Parkinsonia (Gonolkites) convergens'' in the stratigraphic column. The global reference profile for the base of the Bathonian (a GSSP) was ratified as Ravin du Bès, Bas-Auran area, Alpes de Haute Provence, France in 2009. The top of the Bat ...
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Metriorhynchus Geoffroyii
''Metriorhynchus'' is an extinct genus of marine crocodyliform that lived in the oceans during the Late Jurassic. The type species, ''M. brevirostris'' was named in 1829 as a species of ''Steneosaurus'' before being named as a separate genus by the German palaeontologist Christian von Meyer in 1832. The name ''Metriorhynchus'' means "Moderate snout", and is derived from the Greek ''Metrio''- ("moderate") and -''rhynchos'' ("snout"). Discovery and species Fossil specimens referrable to ''Metriorhynchus'' are known from Kimmeridgian (Late Jurassic) deposits of France. Valid species Only one valid species is recognized today, the type species ''M. geoffroyii'' (now called ''M. brevirostris''). ''"Metriorhynchus" hastifer'' and ''"M." palpebrosus'' are generically distinct from the ''Metriorhynchus'' type species, with ''hastifer'' being recovered as a geosaurine. Species in this genus were traditionally classed into two skull groups: longirostrine (long, narrow jaws) and brevirost ...
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Node-based Taxon
Phylogenetic nomenclature is a method of nomenclature for taxa in biology that uses phylogenetic definitions for taxon names as explained below. This contrasts with the traditional approach, in which taxon names are defined by a '' type'', which can be a specimen or a taxon of lower rank, and a description in words. Phylogenetic nomenclature is currently regulated by the '' International Code of Phylogenetic Nomenclature'' (''PhyloCode''). Definitions Phylogenetic nomenclature ties names to clades, groups consisting of an ancestor and all its descendants. These groups can equivalently be called monophyletic. There are slightly different ways of specifying the ancestor, which are discussed below. Once the ancestor is specified, the meaning of the name is fixed: the ancestor and all organisms which are its descendants are included in the named taxon. Listing all these organisms (i.e. providing a full circumscription) requires the full phylogenetic tree to be known. In practice, ther ...
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Torvoneustes Carpenteri
''Torvoneustes'' is an extinct genus of metriorhynchid thalattosuchian. It is known from skull and postcranial remains found in the Kimmeridge Clay Formation of Dorset and Wiltshire, England, and also from Oaxaca, Mexico . The holotype skull of the type species was initially assigned to the species '' Metriorhynchus superciliosus''. Postcranial remains were later discovered from the same quarry as the skull, and then these specimens were recognised as belonging to a new species of '' Dakosaurus'', as ''D. carpenteri''. The species was named to honour Simon Carpenter, an amateur geologist from Frome in Somerset, who discovered the fossils. ''Dakosaurus carpenteri'' was later reassigned to the genus ''Geosaurus'' in 2008. Two years later, it was assigned to its own genus, ''Torvoneustes''. When ''T. carpenteri'' was considered a species of ''Dakosaurus'', its relatively long snout and smaller, more numerous teeth were thought to be features retained from more basal metriorhynch ...
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