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Maniraptoran
Maniraptora is a clade of coelurosaurian dinosaurs which includes the birds and the non-avian dinosaurs that were more closely related to them than to ''Ornithomimus velox''. It contains the major subgroups Avialae, Deinonychosauria, Oviraptorosauria and Therizinosauria. ''Ornitholestes'' and the Alvarezsauroidea are also often included. Together with the next closest sister group, the Ornithomimosauria, Maniraptora comprises the more inclusive clade Maniraptoriformes. Maniraptorans first appear in the fossil record during the Jurassic Period (see ''Eshanosaurus''), and survive today as living birds. Description Maniraptorans are characterized by long arms and three-fingered hands (though reduced or fused in some lineages), as well as a "half-moon shaped" (semi-lunate) bone in the wrist (carpus). In 2004, Tom Holtz and Halszka Osmólska pointed out six other maniraptoran characters relating to specific details of the skeleton. Unlike most other saurischian dinosaurs, which have ...
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Dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is the subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya; their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, having evolved from earlier theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds. Dinosaurs are varied from taxonomic, morphological and ecological standpoints. Birds, at over 10,700 living species, are among ...
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Archaeopteryx
''Archaeopteryx'' (; ), sometimes referred to by its German name, "" ( ''Primeval Bird''), is a genus of bird-like dinosaurs. The name derives from the ancient Greek (''archaīos''), meaning "ancient", and (''ptéryx''), meaning "feather" or "wing". Between the late 19th century and the early 21st century, ''Archaeopteryx'' was generally accepted by palaeontologists and popular reference books as the oldest known bird (member of the group Avialae). Older potential avialans have since been identified, including ''Anchiornis'', ''Xiaotingia'', and ''Aurornis''. ''Archaeopteryx'' lived in the Late Jurassic around 150 million years ago, in what is now southern Germany, during a time when Europe was an archipelago of islands in a shallow warm tropical sea, much closer to the equator than it is now. Similar in size to a Eurasian magpie, with the largest individuals possibly attaining the size of a raven, the largest species of ''Archaeopteryx'' could grow to about in len ...
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Yaverlandia
''Yaverlandia'' (meaning "of Yaverland Point/Yaverland Battery") is a genus of maniraptoran dinosaur. Known from a partial fossil skull (MIWG 1530) found in Lower Cretaceous strata of the Wessex Formation ( Upper Silty Bed; Vectis Formation) on the Isle of Wight. it was described as the earliest known member of the pachycephalosaurid family, but research by Darren Naish shows it to have actually been a theropod, seemingly a maniraptoran. The type species is ''Y. bitholus''.Galton, P.M. (1971) "A primitive dome-headed dinosaur (Ornithischia: Pachycephalosauridae) from the Lower Cretaceous of England and the function of the dome of pachycephalosaurids". ''Journal of Paleontology'', Vol. 45, No. 1, Pages 40–47. Discovery and naming MIWG 1530, the holotype skull, was discovered in 1930, in England and were commented upon by Watson (1930). It was referred to as an iguanodontid of the genus '' Vectisaurus'' in 1936. When Steel (1969) followed Hulke (1879) in listing ''Vectisaurus'' a ...
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Oviraptorosauria
Oviraptorosaurs ("egg thief lizards") are a group of feathered maniraptoran dinosaurs from the Cretaceous Period of what are now Asia and North America. They are distinct for their characteristically short, beaked, parrot-like skulls, with or without bony crests atop the head. They ranged in size from ''Caudipteryx'', which was the size of a turkey, to the 8-meter-long, 1.4-ton ''Gigantoraptor''. The group (along with all maniraptoran dinosaurs) is close to the ancestry of birds. Some researchers such as Maryanska ''et al'' (2002) and Osmólska ''et al.'' (2004) have proposed that they may represent primitive flightless birds.Osmólska, Halszka, Currie, Philip J., Brasbold, Rinchen (2004) "The Dinosauria" Weishampel, Dodson, Osmólska. "Chapter 8 Oviraptorosauria" University of California Press. The most complete oviraptorosaur specimens have been found in Asia. The North American oviraptorosaur record is sparse.Varricchio, D. J. 2001. Late Cretaceous oviraptorosaur (Theropoda) ...
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Therizinosauria
Therizinosaurs (once called segnosaurs) were large herbivorous theropod dinosaurs whose fossils have been found across the Early to Late Cretaceous deposits in Asia and North America. Various features of the forelimbs, skull and pelvis unite these finds as both theropods and maniraptorans, close relatives to birds. The name of the representative genus, ''Therizinosaurus'', is derived from the Greek (, 'to reap' or 'scythe')Translated paper
and (, 'lizard'). The older representative, ''Segnosaurus'', is derived from the ('slow') and the Greek .


History of research


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Elopteryx
''Elopteryx'' is a genus of, perhaps troodontid, maniraptoran theropod dinosaur based on fragmentary fossils found in late Cretaceous Period rocks of Romania. The single species, ''Elopteryx nopcsai'', is known only from very incomplete material, and therefore is considered a '' nomen dubium'' ("dubious name") by most paleontologists. History of discovery and naming Initial finds In the late nineteenth or early twentieth century, the famous Hungarian paleontologist Franz Nopcsa von Felső-Szilvás found near Sînpetru, in what is now the Romanian region of Transylvania, some bone fragments of a small theropod. These were acquired by the British Museum of Natural History. In 1913, curator Charles William Andrews named these as the type species ''Elopteryx nopcsai''. The genus name ''Elopteryx'' is from Ancient Greek ''helos'' (ἕλος), "marsh" + ''pteryx'' (πτέρυξ), "wing". The specific name honors Nopcsa. Initially, ''Elopteryx'' was described from its holotype, a pro ...
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Fukuivenator
''Fukuivenator'' ("hunter of Fukui Prefecture") is an extinct genus of therizinosaurian theropod dinosaur from the Early Cretaceous of Japan. Discovery and naming The type species ''Fukuivenator paradoxus'' was in 2016 named and described by Yoichi Azuma, Xu Xing, Masateru Shibata, Soichiro Kawabe, Kazunori Miyata and Takuya Imai. The generic name combines a reference to the Fukui prefecture with Latin ''venator'', "hunter". The specific name refers to the paradoxical combination of traits shown by the species. The rocks in which the skeleton of ''Fukuivenator'', holotype FPDM-V8461, was found in August 2007 belong to the Kitadani Formation, which is probably of Barremian or Aptian age. Radiometric dating of nearby rock units has given this formation an estimated age of somewhere between 127 and 115 million years old. The holotype consists of a partial skeleton with skull. The skeleton of ''F. paradoxus'' is currently the most complete non-avian dinosaur fossil found in Ja ...
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Pedopenna
''Pedopenna'' (meaning "foot feather") is a genus of small, feathered, maniraptoran dinosaur from the Daohugou Beds in China. It is possibly older than ''Archaeopteryx'', though the age of the Daohugou Beds where it was found is debated. A majority of studies suggest that beds probably date from between the late Middle Jurassic (168 million years ago) and early Late Jurassic Period (164-152 million years ago). The name ''Pedopenna'' refers to the long pennaceous feathers on the metatarsus; ''daohugouensis'' refers to the locality of Daohugou, where the holotype was found. ''Pedopenna daohugouensis'' probably measured 1 meter (3 ft) or less in length, but since this species is only known from the hind legs, the actual length is difficult to estimate. ''Pedopenna'' was originally classified as a paravian, the group of maniraptoran dinosaurs that includes both deinonychosaurs and avialans (the lineage including modern birds), but some scientists have classified it as a tru ...
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Kakuru
''Kakuru'' is a genus of theropod dinosaur from the early Cretaceous Period. The only described species, ''Kakuru kujani'', is known primarily from evidence of a single tibia, which had been fossilised through a rare process in which the bone through hydration turned to opal. The bone was dug up at the opal fields of Andamooka, South Australia. The opalised tibia was exhibited by a gem shop in 1973 and by chance brought to the attention of paleontologist Neville Pledge. The owner at the time, a certain A. Fleming, allowed pictures and two casts to be made but eventually the specimen was sold at an auction to an anonymous buyer. It was presumed lost to science. In 2004, however, the South Australian Museum succeeded in procuring the fossil for $22,000. ''Kakuru'' was formally named in 1980 by Pledge and Ralph Molnar.R. E. Molnar and N. S. Pledge, 1980, "A new theropod dinosaur from South Australia", ''Alcheringa'' 4: 281-287 The type species is ''Kakuru kujani''. The generic na ...
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Coelurosauria
Coelurosauria (; from Greek, meaning "hollow tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyrannosaurs, ornithomimosaurs, and maniraptorans; Maniraptora includes birds, the only known dinosaur group alive today. Most feathered dinosaurs discovered so far have been coelurosaurs. Philip J. Currie had considered it likely and probable that all coelurosaurs were feathered. However, several skin impressions found for some members of this group show pebbly, scaly skin, indicating that feathers did not completely replace scales in all taxa. In the past, Coelurosauria was used to refer to all small theropods, but this classification has since been abolished. Anatomy Bodyplan The studying of anatomical traits in coelurosaurs indicates that the last common ancestor had evolved the ability to eat and digest plant matter, adapting to an omniv ...
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Bird
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweight skeleton. Birds live worldwide and range in size from the bee hummingbird to the ostrich. There are about ten thousand living species, more than half of which are passerine, or "perching" birds. Birds have whose development varies according to species; the only known groups without wings are the extinct moa and elephant birds. Wings, which are modified forelimbs, gave birds the ability to fly, although further evolution has led to the loss of flight in some birds, including ratites, penguins, and diverse endemic island species. The digestive and respiratory systems of birds are also uniquely adapted for flight. Some bird species of aquatic environments, particularly seabirds and some waterbirds, have further evolved for swimming. B ...
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Middle Jurassic
The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations containing land animal fossils include the Forest Marble Formation in England, the Kilmaluag Formation in Scotland,British Geological Survey. 2011Stratigraphic framework for the Middle Jurassic strata of Great Britain and the adjoining continental shelf: research report RR/11/06 British Geological Survey, Keyworth, Nottingham. the Daohugou Beds in China, the Itat Formation in Russia, and the Isalo III Formation of western Madagascar. Paleogeography During the Middle Jurassic Epoch, Pangaea began to separate into Laurasia and Gondwana, and the Atlantic Ocean formed. Eastern Laurasia was tectonically active as the Cimmerian plate continued to collide with Laurasia's southern coast, completely closing the Paleo-Tethys Ocean. A subduction zone ...
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