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Ichthyosauriformes
The Ichthyosauriformes are a group of marine reptiles, belonging to the Ichthyosauromorpha, that lived during the Mesozoic. The stem clade Ichthyosauriformes was in 2014 defined by Ryosuke Motani and colleagues as the group consisting of all ichthyosauromorphs that are more closely related to '' Ichthyosaurus communis'' than to '' Hupehsuchus nanchangensis''. Their synapomorphies include the possession of a long nasal bone, stretching to the front beyond the nostril; large scleral rings, filling the eye sockets; a narrow snout in top view; and converging digits with little space between them. The Ichthyosauriformes probably split off in the Early Triassic, about 250 million years ago; the last known forms lived in the middle Cretaceous. A basal ichthyosauriform is ''Cartorhynchus''; more derived species are part of the Ichthyopterygia which again include the Ichthyosauria Ichthyosaurs (Ancient Greek for "fish lizard" – and ) are large extinct marine reptiles. Ichthyosa ...
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Ichthyosauromorpha
The Ichthyosauromorpha are an extinct clade of marine reptiles consisting of the Ichthyosauriformes and the Hupehsuchia, living during the Mesozoic. The node clade Ichthyosauromorpha was first defined by Ryosuke Motani ''et al.'' in 2014 as the group consisting of the last common ancestor of ''Ichthyosaurus communis'' and ''Hupehsuchus nanchangensis'', and all its descendants. Their synapomorphies, unique derived traits, include: the presence of an anterior flange on the humerus and Radius (bone), radius; the lower end of the ulna being as wide as or wider than the upper end, the forelimb being as long as or longer than the hindlimb, the hand having at least three quarters of the length of the upper arm and lower arm combined, the fibula extending behind the level of the thighbone, and the transverse process of the vertebral neural arch being reduced or absent. The Ichthyosauromorpha probably originated in China during the upper Lower Triassic period, about 248 million years ag ...
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Cartorhynchus
''Cartorhynchus'' (meaning "shortened snout") is an extinct genus of early ichthyosauriform marine reptile that lived during the Early Triassic epoch, about 248 million years ago. The genus contains a single species, ''Cartorhynchus lenticarpus'', named in 2014 by Ryosuke Motani and colleagues from a single nearly-complete skeleton found near Chaohu, Anhui Province, China. Along with its close relative '' Sclerocormus'', ''Cartorhynchus'' was part of a diversification of marine reptiles that occurred suddenly (over about one million years) during the Spathian substage, soon after the devastating Permian-Triassic extinction event, but they were subsequently driven to extinction by volcanism and sea level changes by the Middle Triassic. Measuring about long, ''Cartorhynchus'' was a small animal with a lizard-like body and a short torso; it probably swam in an eel-like manner at slow speeds. Its limbs bore extensive cartilage and could bend like flippers, which may have allo ...
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Sclerocormus
''Sclerocormus'' is an extinct genus of ichthyosauriform from the early Triassic period. The fossil was discovered in the central Anhui Province, China. It is currently only known from one specimen, however the fossil is mostly complete and as such further increases the understanding of the early evolution of ichthyosaurs. Description ''Sclerocormus'' is much larger than its closest relative ''Cartorhynchus'', with a total body length of and body mass of more than . Its proportions were unusual amongst basal ichthyosauriformes, with a short, heavily built trunk, a very long tail over 92 cm long (58% of the total body length), and a small skull with a short, narrow snout and toothless jaws. Like ''Cartorhynchus'', the skull of ''Sclerocormus'' is wide, with a short, edentulous snout much narrower than the skull roof. The skull is unusually short at only 6.25% of its body length, compared to 12% in ''Chaohusaurus'' and 15% in ''Hupehsuchus''. Unusually, the nasals of ''Sc ...
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Ichthyosauria
Ichthyosaurs (Ancient Greek for "fish lizard" – and ) are large extinct marine reptiles. Ichthyosaurs belong to the order known as Ichthyosauria or Ichthyopterygia ('fish flippers' – a designation introduced by Sir Richard Owen in 1842, although the term is now used more for the parent clade of the Ichthyosauria). Ichthyosaurs thrived during much of the Mesozoic era; based on fossil evidence, they first appeared around 250 million years ago ( Ma) and at least one species survived until about 90 million years ago, into the Late Cretaceous. During the Early Triassic epoch, ichthyosaurs and other ichthyosauromorphs evolved from a group of unidentified land reptiles that returned to the sea, in a development similar to how the mammalian land-dwelling ancestors of modern-day dolphins and whales returned to the sea millions of years later, which they gradually came to resemble in a case of convergent evolution. Ichthyosaurs were particularly abundant in the Late Triassic a ...
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Early Triassic
The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between Ma and Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series, which is a unit in chronostratigraphy. The Early Triassic is the oldest epoch of the Mesozoic Era. It is preceded by the Lopingian Epoch (late Permian, Paleozoic Era) and followed by the Middle Triassic Epoch. The Early Triassic is divided into the Induan and Olenekian ages. The Induan is subdivided into the Griesbachian and Dienerian subages and the Olenekian is subdivided into the Smithian and Spathian subages. The Lower Triassic series is coeval with the Scythian Stage, which is today not included in the official timescales but can be found in older literature. In Europe, most of the Lower Triassic is composed of Buntsandstein, a lithostratigraphic unit of continental red beds. The Early Triassic and partly also the Middle Triassic span the in ...
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Synapomorphies
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis T ...
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Olenekian First Appearances
In the geologic timescale, the Olenekian is an age in the Early Triassic epoch; in chronostratigraphy, it is a stage in the Lower Triassic series. It spans the time between Ma and Ma (million years ago). The Olenekian is sometimes divided into the Smithian and the Spathian subages or substages. The Olenekian follows the Induan and is followed by the Anisian (Middle Triassic). The Olenekian saw the deposition of a large part of the Buntsandstein in Europe. The Olenekian is roughly coeval with the regional Yongningzhenian Stage used in China. Stratigraphic definitions The Olenekian Stage was introduced into scientific literature by Russian stratigraphers in 1956. The stage is named after Olenëk in Siberia. Before the subdivision in Olenekian and Induan became established, both stages formed the Scythian Stage, which has since disappeared from the official timescale. The base of the Olenekian is at the lowest occurrence of the ammonoids '' Hedenstroemia'' or '' Meekoceras gra ...
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Mesozoic Reptiles
The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Cretaceous Periods. It is characterized by the dominance of archosaurian reptiles, like the dinosaurs; an abundance of conifers and ferns; a hot greenhouse climate; and the tectonic break-up of Pangaea. The Mesozoic is the middle of the three eras since complex life evolved: the Paleozoic, the Mesozoic, and the Cenozoic. The era began in the wake of the Permian–Triassic extinction event, the largest well-documented mass extinction in Earth's history, and ended with the Cretaceous–Paleogene extinction event, another mass extinction whose victims included the non-avian dinosaurs, pterosaurs, mosasaurs, and plesiosaurs. The Mesozoic was a time of significant tectonic, climatic, and evolutionary activity. The era witnessed the gradual rifting ...
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Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of the entire Phanerozoic. The name is derived from the Latin ''creta'', "chalk", which is abundant in the latter half of the period. It is usually abbreviated K, for its German translation ''Kreide''. The Cretaceous was a period with a relatively warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now- extinct marine reptiles, ammonites, and rudists, while dinosaurs continued to dominate on land. The world was ice free, and forests extended to the poles. During this time, new groups of mammals and birds appeared. During the Early Cretaceous, flowering plants appeared and began to rapidly diversify, becoming the dominant group of plants across the Earth b ...
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Scleral Ring
Sclerotic rings are rings of bone found in the eyes of many animals in several groups of vertebrates, except for mammals and crocodilians. They can be made up of single bones or multiple segments and take their name from the sclera. They are believed to have a role in supporting the eye, especially in animals whose eyes are not spherical, or which live underwater. Fossil sclerotic rings are known for a variety of extinct animals, including ichthyosaurs, pterosaurs, and dinosaurs, but are often not preserved. File:Tawny Frogmouth Skull.jpg, A skull of an extant tawny frogmouth, showing large sclerotic rings. File:Uroplatus_phantasticus_skull1.jpg, A skull of an extant satanic leaf-tailed gecko, showing large sclerotic rings. File:Viatkogorgon skull.png, Skull and sclerotic ring of the gorgonopsian ''Viatkogorgon''. File:Prosaurolophus maximus3.JPG, Sclerotic ring of the hadrosaur ''Prosaurolophus ''Prosaurolophus'' (; meaning "before ''Saurolophus''", in comparison to the late ...
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Nasal Bone
The nasal bones are two small oblong bones, varying in size and form in different individuals; they are placed side by side at the middle and upper part of the face and by their junction, form the bridge of the upper one third of the nose. Each has two surfaces and four borders. Structure The two nasal bones are joined at the midline internasal suture and make up the bridge of the nose. Surfaces The ''outer surface'' is concavo-convex from above downward, convex from side to side; it is covered by the procerus and nasalis muscles, and perforated about its center by a foramen, for the transmission of a small vein. The ''inner surface'' is concave from side to side, and is traversed from above downward, by a groove for the passage of a branch of the nasociliary nerve. Articulations The nasal articulates with four bones: two of the cranium, the frontal and ethmoid, and two of the face, the opposite nasal and the maxilla. Other animals In primitive bony fish and tetrapod ...
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Ichthyosaurus Communis
''Ichthyosaurus'' (derived from Greek ' () meaning 'fish' and ' () meaning 'lizard') is a genus of ichthyosaurs from the Early Jurassic (Hettangian - Pliensbachian), with possible Late Triassic record, from Europe (Belgium, England, Germany, Switzerland, and Portugal). It is among the best known ichthyosaur genera, as it is the type genus of the order Ichthyosauria.Maisch MW, Matzke AT. 2000. The Ichthyosauria. ''Stuttgarter Beiträge zur Naturkunde, Serie B (Geologie und Paläontologie)'' 298: 1-159McGowan C, Motani R. 2003. Ichthyopterygia. – In: Sues, H.-D. (ed.): ''Handbook of Paleoherpetology, Part 8, Verlag Dr. Friedrich Pfeil'', 175 pp., 101 figs., 19 plts; MünchenMaisch MW, Reisdorf AG, Schlatter R, Wetzel A. 2008. A large skull of Ichthyosaurus (Reptilia: Ichthyosauria) from the Lower Sinemurian (Lower Jurassic) of Frick (NW Switzerland). ''Swiss Journal of Geosciences'' 101: 617-627. History of discovery ''Ichthyosaurus'' was the first complete fossil to be discovere ...
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