Evolution Of Seed Size
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Evolution Of Seed Size
The first seeded plants emerged in the late Devonian 370 million years ago. Selection pressures shaping seed size stem from physical and biological sources including drought, predation, seedling-seedling competition, optimal dormancy depth, and dispersal. History Since the evolution of the first seeded plants ~370 million years ago, the largest change in seed size was found to be at the divergence of gymnosperms and angiosperms ~325 million years ago, but overall, the divergence of seed size appears to take place relatively consistently through evolutionary time. Seed mass has been found to be phylogenetically conservative with most differences in mean seed mass within types of seed dispersal (dispersal modes) being phylogenetic. This type of information gives us clues about how seed size evolved. Dating fossilized seeds of various sizes and comparing them with the presence of possible animal dispersers and the environmental conditions of the time is another technique used to stu ...
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Devonian
The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, where rocks from this period were first studied. The first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents. By the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. The arthropod groups of myriapods, arachnids and hexapods also became well-established early in this period, after starting their expansion to land at least from the Ordovician period. Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fishes. The placoderms began dominating ...
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Endosperm
The endosperm is a tissue produced inside the seeds of most of the flowering plants following double fertilization. It is triploid (meaning three chromosome sets per nucleus) in most species, which may be auxin-driven. It surrounds the embryo and provides nutrition in the form of starch, though it can also contain oils and protein. This can make endosperm a source of nutrition in animal diet. For example, wheat endosperm is ground into flour for bread (the rest of the grain is included as well in whole wheat flour), while barley endosperm is the main source of sugars for beer production. Other examples of endosperm that forms the bulk of the edible portion are coconut "meat" and coconut "water", and corn. Some plants, such as orchids, lack endosperm in their seeds. Origin of endosperm Ancestral flowering plants have seeds with small embryos and abundant endosperm. The evolutionary development of flowering plants trends towards plants with mature seeds with little or no endosp ...
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Evolution Of Plants
The evolution of plants has resulted in a wide range of complexity, from the earliest algal mats, through multicellular marine and freshwater green algae, terrestrial bryophytes, lycopods and ferns, to the complex gymnosperms and angiosperms (flowering plants) of today. While many of the earliest groups continue to thrive, as exemplified by red and green algae in marine environments, more recently derived groups have displaced previously ecologically dominant ones; for example, the ascendance of flowering plants over gymnosperms in terrestrial environments. There is evidence that cyanobacteria and multicellular photosynthetic eukaryotes lived in freshwater communities on land as early as 1 billion years ago, and that communities of complex, multicellular photosynthesizing organisms existed on land in the late Precambrian, around . Evidence of the emergence of embryophyte land plants first occurs in the mid-Ordovician (~), and by the middle of the Devonian (~), many of the featur ...
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Seed Dispersal Syndrome
Seed dispersal syndromes are morphological characters of seeds correlated to particular seed dispersal agents.Clobert, J., Le Galliard, J.F., Cote, J., Meylan, S. & Massot, M. (2009). Informed dispersal, heterogeneity in animal dispersal syndromes and the dynamics of spatially structured populations. Ecology Letters, 12, 197-209.Griz, L.M.S. & Machado, I.C.S. (2001). Fruiting phenology and seed dispersal syndromes in caatinga, a tropical dry forest in the northeast of Brazil. Journal of Tropical Ecology, 17, 303-321.Link, A. & Stevenson, P.R. (2004). Fruit dispersal syndromes in animal disseminated plants at Tinigua National Park, Colombia. Revista Chilena de Historia Natural, 77, 319-334.Tamboia, T., Cipollini, M.L. & Levey, D.J. (1996). An Evaluation of Vertebrates Seed Dispersal Syndromes in Four Species of Black Nightshade. Oecologia, 107(4), 522-532. Dispersal is the event by which individuals move from the site of their parents to establish in a new area.Herrera, C. M. & Pellmyr ...
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Cross Pollinate
Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, birds, and bats; water; wind; and even plants themselves, when self-pollination occurs within a closed flower. Pollination often occurs within a species. When pollination occurs between species, it can produce hybrid offspring in nature and in plant breeding work. In angiosperms, after the pollen grain (gametophyte) has landed on the stigma, it germinates and develops a pollen tube which grows down the style until it reaches an ovary. Its two gametes travel down the tube to where the gametophyte(s) containing the female gametes are held within the carpel. After entering an ovum cell through the micropyle, one male nucleus fuses with the polar bodies to produce the endosperm tissues, while the other fuses with the ovule to produce the embryo. Hen ...
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Seed Predation
Seed predation, often referred to as granivory, is a type of plant-animal interaction in which granivores (seed predators) feed on the seeds of plants as a main or exclusive food source,Hulme, P.E. and Benkman, C.W. (2002) "Granivory", pp. 132–154 in ''Plant animal Interactions: An Evolutionary Approach'', ed. C.M. Herrera and O. Pellmyr. Oxford: Blackwell. . in many cases leaving the seeds damaged and not viable. Granivores are found across many families of vertebrates (especially mammals and birds) as well as invertebrates (mainly insects); thus, seed predation occurs in virtually all terrestrial ecosystems. Seed predation is commonly divided into two distinctive temporal categories, pre-dispersal and post-dispersal predation, which affect the fitness of the parental plant and the dispersed offspring (the seed), respectively. Mitigating pre- and post-dispersal predation may involve different strategies. To counter seed predation, plants have evolved both physical defenses ( ...
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Bet Hedging (biology)
Biological bet hedging occurs when organisms suffer decreased fitness in their typical conditions in exchange for increased fitness in stressful conditions. Biological bet hedging was originally proposed to explain the observation of a Soil seed bank, seed bank, or a reservoir of ungerminated seeds in the soil. For example, an Annual plant, annual plant's Fitness (biology), fitness is maximized for that year if all of its seeds germinate. However, if a drought occurs that kills germinated plants, but not ungerminated seeds, plants with seeds remaining in the seed bank will have a fitness advantage. Therefore, it can be advantageous for plants to "hedge their bets" in case of a drought by producing some seeds that germinate immediately and other seeds that lie dormant. Other examples of biological bet hedging include Polyandry in nature, female multiple mating, foraging behavior in Bumblebee, bumble bees, nutrient storage in rhizobia, and Multidrug tolerance, bacterial persistence in ...
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Butternut (tree)
''Juglans cinerea'', commonly known as butternut or white walnut,Snow, Charles Henry ''The Principal Species of Wood: Their Characteristic Properties'' 2nd ed. New York: John Wiley & Sons, 1908. p. 56. is a species of walnut native to the eastern United States and southeast Canada. Distribution The distribution range of ''J. cinerea'' extends east to New Brunswick, and from southern Quebec west to Minnesota, south to northern Alabama and southwest to northern Arkansas. It is absent from most of the Southern United States. The species also proliferates at middle elevations (about above sea level) in the Columbia River basin, Pacific Northwest; as an off-site species. Trees with (over mature) class range diameter at breast height were noted in the Imnaha River drainage as late as January 26, 2015. Butternut favors a cooler climate than black walnut and its range does not extend into the Deep South. Its northern range extends into Wisconsin and Minnesota where the growing s ...
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Pecan
The pecan (''Carya illinoinensis'') is a species of hickory native to the southern United States and northern Mexico in the region of the Mississippi River. The tree is cultivated for its seed in the southern United States, primarily in Georgia, New Mexico, Texas, and Mexico, which produces nearly half of the world total. The seed is an edible nut used as a snack and in various recipes, such as praline candy and pecan pie. The pecan is the state nut of Alabama, Arkansas, California, and Texas and is also the state tree of Texas. Name "Pecan" is from an Algonquin word, variously referring to pecans, walnuts, and hickory nuts. There are many pronunciations, some regional and others not.See "Pecan" at Wiktionary. The most common American pronunciation is . There is little agreement in the United States regarding the "correct" pronunciation, even regionally. Growth The pecan tree is a large deciduous tree, growing to in height, rarely to .Flora of North America''Carya illinoi ...
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Hickory
Hickory is a common name for trees composing the genus ''Carya'', which includes around 18 species. Five or six species are native to China, Indochina, and India (Assam), as many as twelve are native to the United States, four are found in Mexico, and two to four are native to Canada. A number of hickory species are used for products like edible nuts or wood. Hickories are temperate forest trees with pinnately compound leaves and large nuts. Hickory flowers are small, yellow-green catkins produced in spring. They are wind-pollinated and self-incompatible. The fruit is a globose or oval nut, long and diameter, enclosed in a four-valved husk, which splits open at maturity. The nut shell is thick and bony in most species, and thin in a few, notably the pecan (''C. illinoinensis''); it is divided into two halves, which split apart when the seed germinates. Etymology The name "hickory" derives from a Native American word in an Algonquian language (perhaps Powhatan). It is a ...
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Old-growth Forest
An old-growth forestalso termed primary forest, virgin forest, late seral forest, primeval forest, or first-growth forestis a forest that has attained great age without significant disturbance, and thereby exhibits unique ecological features, and might be classified as a climax community. The Food and Agriculture Organization of the United Nations defines primary forests as naturally regenerated forests of native tree species where there are no clearly visible indications of human activity and the ecological processes are not significantly disturbed. More than one-third (34 percent) of the world's forests are primary forests. Old-growth features include diverse tree-related structures that provide diverse wildlife habitat that increases the biodiversity of the forested ecosystem. Virgin or first-growth forests are old-growth forests that have never been logged. The concept of diverse tree structure includes multi-layered canopies and canopy gaps, greatly varying tree height ...
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Pliocene
The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58See the 2014 version of the ICS geologic time scale
million years ago. It is the second and most recent epoch of the Neogene Period in the . The Pliocene follows the Epoch and is followed by the Epoch. Prior to the 2009 ...
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