Cynodonts
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Cynodonts
The cynodonts () (clade Cynodontia) are a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Cynodonts had a wide variety of lifestyles, including carnivory and herbivory. Mammals are cynodonts, as are their extinct ancestors and close relatives, having evolved from advanced probainognathian cynodonts during the Late Triassic. All other cynodont lines went extinct, with the last known non-mammalian cynodont group, the Tritylodontidae, having its youngest records in the Early Cretaceous. Description Early cynodonts have many of the skeletal characteristics of mammals. The teeth were fully differentiated and the braincase bulged at the back of the head. Outside of some crown-group mammals (notably the therians), all cynodonts probably laid eggs. The temporal fenestrae were much larger than those of their ancestors, and the widening of the zygomatic arch in ...
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Therapsid
Therapsida is a major group of eupelycosaurian synapsids that includes mammals, their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more underneath the body, as opposed to the sprawling posture of many reptiles and salamanders. Therapsids evolved from "pelycosaurs", specifically within the Sphenacodontia, more than 279.5 million years ago. They replaced the "pelycosaurs" as the dominant large land animals in the Middle Permian through to the Early Triassic. In the aftermath of the Permian–Triassic extinction event, therapsids declined in relative importance to the rapidly diversifying reptiles during the Middle Triassic. The therapsids include the cynodonts, the group that gave rise to mammals ( Mammaliaformes) in the Late Triassic, around 225 million years ago. Of the non-mammalian therapsids, only cynodonts survived beyond the end of the Triassic, with the only other ...
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Brasilitherium Riograndensis
''Brasilodon'' ("tooth from Brazil") is an extinct genus of small, mammal-like cynodonts that lived in what is now Brazil during the Norian age of the Late Triassic epoch, about 225.42 million years ago. While no complete skeletons have been found, the length of ''Brasilodon'' has been estimated at around . Its dentition shows that it was most likely an insectivore. The genus is monotypic, containing only the species ''B. quadrangularis''. ''Brasilodon'' belongs to the family Brasilodontidae, whose members were some of the closest relatives of mammals, the only cynodonts alive today. Two other brasilodontid genera, ''Brasilitherium'' and ''Minicynodon'', are now considered to be junior synonyms of ''Brasilodon''. Discovery and naming The first three specimens referred to ''Brasilodon quadrangularis'' were found at the Linha São Luiz site, a quarry near the town of Faxinal do Soturno in the state of Rio Grande do Sul. The rocks where ''Brasilodon'' was found belong to t ...
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Eutheriodont
Eutheriodontia is a clade of therapsids which appear during the Middle Permian and which includes therocephalians and cynodonts, this latter group including mammals and related forms. With the dicynodonts, they form one of two lineages of therapsids that survived the Permian extinction and which diversified again during the Triassic, before the majority of them disappeared before or during the Triassic–Jurassic extinction, except for a lineage of cynodonts that would later give rise to mammals. Classification The clade was named in 1986 by James Allen Hopson and Herbert Richard Barghusen, the name meaning the "true Theriodontia". Within Hopson's system, the Eutheriodontia are the sister group of the Gorgonopsia within the Theriodontia. A close relationship between therocephalians and cynodonts had been recognized for many years. In 2001 the Eutheriodontia were defined as the least inclusive clade including Mammalia and ''Bauria''.James A. Hopson and James W. Kitching, 20 ...
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Epicynodontia
Epicynodontia is a clade of cynodont therapsids that includes most cynodonts, such as galesaurids, thrinaxodontids, and Eucynodontia (including mammals). It was erected as a stem-based taxon by Hopson and Kitching (2001) and defined as the most inclusive clade containing Mammalia and excluding ''Procynosuchus'', a Late Permian genus that is one of the most basal cynodonts. Below is a cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ... from Ruta, Botha-Brink, Mitchell and Benton (2013) showing one hypothesis of cynodont relationships: References Cynodonts Lopingian first appearances Extant Permian first appearances Tetrapod unranked clades Taxa named by James Hopson Taxa named by James Kitching {{Therapsid-stub ...
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Probainognathia
Probainognathia is one of the two major subgroups of the clade Eucynodontia, the other being Cynognathia. The earliest forms were carnivorous and insectivorous, though some groups eventually also evolved herbivorous diets. The earliest and most basal probainognathian is the Middle Triassic (Anisian) aged ''Lumkuia'', from South Africa, though probainognathians would not become prominent until the mid Norian stage of the Late Triassic. Three groups survived the extinction at the end of Triassic: Tritheledontidae and Tritylodontidae, which both survived until the Jurassic—the latter even into the Cretaceous (''Montirictus'' and '' Xenocretosuchus'')—and Mammaliaformes, which includes the mammals. Phylogeny Below is a cladogram from Ruta, Botha-Brink, Mitchell and Benton (2013) showing one hypothesis of cynodont relationships: See also * Evolution of mammals * List of prehistoric mammals This is an incomplete list of prehistoric mammals. It does not include extant mam ...
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Tritylodontidae
Tritylodontidae ("three-knob teeth", named after the shape of their cheek teeth) is an extinct family of small to medium-sized, highly specialized mammal-like cynodonts, bearing several mammalian traits like erect limbs, endothermy and details of the skeleton. They were the last-known family of the non-mammaliaform synapsids, persisting into the Early Cretaceous. Most tritylodontids are thought to have been herbivorous, feeding on vegetation, such as stems, leaves, and roots, although at least one may have had a more omnivorous diet. Tritylodontid fossils are found in the Americas, South Africa, and Eurasia—they appear to have had an almost global distribution, including Antarctica. Description The skull of tritylodontids had a high sagittal crest. They retained the primitive condition of the joint between the quadrate bone of the skull and the articular bone of the lower jaw—the retention of the joint is one of the reasons they are technically regarded to not be mam ...
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Early Cretaceous
The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian-Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining early Aptian Oceanic Anoxic Event 1a (OAE1a) was carbon isotope dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java-Manihiki-Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. The Ontong Java Plateau today covers an area of 1,860,000 km2. In the Indian Ocean another LIP began to form at c. 120 Ma, the Kerguelen P ...
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Permian–Triassic Extinction Event
The Permian–Triassic (P–T, P–Tr) extinction event, also known as the Latest Permian extinction event, the End-Permian Extinction and colloquially as the Great Dying, formed the boundary between the Permian and Triassic geologic periods, as well as between the Paleozoic and Mesozoic eras, approximately 251.9 million years ago. It is the Earth's most severe known extinction event, with the extinction of 57% of biological families, 83% of genera, 81% of marine species and 70% of terrestrial vertebrate species. It was the largest known mass extinction of insects. There is evidence for one to three distinct pulses, or phases, of extinction. The scientific consensus is that the main cause of extinction was the large amount of carbon dioxide emitted by the volcanic eruptions that created the Siberian Traps, which elevated global temperatures, and in the oceans led to widespread anoxia and acidification. Proposed contributing factors include: the emission of much addition ...
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Mammal
Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or hair, and three middle ear bones. These characteristics distinguish them from reptiles (including birds) from which they diverged in the Carboniferous, over 300 million years ago. Around 6,400 extant species of mammals have been described divided into 29 orders. The largest orders, in terms of number of species, are the rodents, bats, and Eulipotyphla (hedgehogs, moles, shrews, and others). The next three are the Primates (including humans, apes, monkeys, and others), the Artiodactyla ( cetaceans and even-toed ungulates), and the Carnivora (cats, dogs, seals, and others). In terms of cladistics, which reflects evolutionary history, mammals are the only living members of the Synapsida (synapsids); this clade, together with Saur ...
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Mammal
Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or hair, and three middle ear bones. These characteristics distinguish them from reptiles (including birds) from which they diverged in the Carboniferous, over 300 million years ago. Around 6,400 extant species of mammals have been described divided into 29 orders. The largest orders, in terms of number of species, are the rodents, bats, and Eulipotyphla (hedgehogs, moles, shrews, and others). The next three are the Primates (including humans, apes, monkeys, and others), the Artiodactyla ( cetaceans and even-toed ungulates), and the Carnivora (cats, dogs, seals, and others). In terms of cladistics, which reflects evolutionary history, mammals are the only living members of the Synapsida (synapsids); this clade, together with Saur ...
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Bienotherium Yunnanense
''Bienotherium'' is an extinct genus of cynodonts from the Early Jurassic of China discovered by Bian Meinian (Mei Nien Bien). Despite its size, it is closely related to '' Lufengia'', and is the largest tritylodont from the Lufeng Formation in China. ''Bienotherium'' had four incisors, no canines, and back molar-like teeth, which it used to chew tough plant material. Description ''Bienotherium'' is defined as being big and robust compared to other tritylodonts, and also by exposed maxillaries in the skull, an unusually long diastema and thin zygomatic bone In the human skull, the zygomatic bone (from grc, ζῠγόν, zugón, yoke), also called cheekbone or malar bone, is a paired irregular bone which articulates with the maxilla, the temporal bone, the sphenoid bone and the frontal bone. It is s .... References Further reading
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Crown Group
In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. It is thus a way of defining a clade, a group consisting of a species and all its extant or extinct descendants. For example, Neornithes (birds) can be defined as a crown group, which includes the most recent common ancestor of all modern birds, and all of its extant or extinct descendants. The concept was developed by Willi Hennig, the formulator of phylogenetic systematics, as a way of classifying living organisms relative to their extinct relatives in his "Die Stammesgeschichte der Insekten", and the "crown" and "stem" group terminology was coined by R. P. S. Jefferies in 1979. Though formulated in the 1970s, the term was not commonly used until its reintroduction in 2000 by Graham Budd and Sören Jensen. Contents of the crown gr ...
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