Coccosteid
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Coccosteid
Coccosteidae is a family of arthrodire placoderms from the Early to Late Devonian. Fossils appear in various strata in Europe, North America and China. Phylogeny Coccosteidae belongs to the larger clade Coccosteomorphi, which together with its sister clade Pachyosteomorphi forms the group Eubrachythoraci. The phylogeny of Coccosteidae can be shown in the cladogram below: Genera '' Belgiosteus'' A genus of very large coccosteids. Species are found in Middle Devonian Belgium and China. '' Clarkosteus'' ''Coccosteus'' The type genus of the family. Numerous species are found in Middle to Upper Devonian strata throughout Europe and parts of North America. ''Dickosteus'' ''Jiuchengia'' The earliest known coccosteid from Late Emsian Yunnan province, China. It is distinguished from other coccosteids by having an elongated occipital. '' Livosteus'' A genus of very large coccosteids known from Middle to Late Devonian strata of Eastern Europe. ''Millerosteus'' A genus of very small ...
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Protitanichthys Rockportensis
''Protitanichthys'' is an extinct genus of comparatively large coccosteid arthrodire placoderms from the Middle Devonian of the eastern United States. Fossils are found primarily in the Eifelian-epoch aged Delaware Limestone of Ohio, and the Lower Givetian-aged Rockport Quarry Limestone of Michigan Description ''Protitanichthys'' is very similar to other coccosteids, though the skull is proportionally narrower, and the orbits are comparatively smaller. As mentioned earlier, species are quite large for coccosteids, surpassed only in size by the Old World genus, '' Livosteus''. Phylogeny ''Protitanichthys'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny of ''Protitanichthys'': Species ''P. fossatus'' The type species of the genus. The description of ''P. fossatus'' is based on a cranial roof, possibly long, found in the Eifelian-aged Delaware ...
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Protitanichthys
''Protitanichthys'' is an extinct genus of comparatively large coccosteid arthrodire placoderms from the Middle Devonian of the eastern United States. Fossils are found primarily in the Eifelian-epoch aged Delaware Limestone of Ohio, and the Lower Givetian-aged Rockport Quarry Limestone of Michigan Description ''Protitanichthys'' is very similar to other coccosteids, though the skull is proportionally narrower, and the orbits are comparatively smaller. As mentioned earlier, species are quite large for coccosteids, surpassed only in size by the Old World genus, '' Livosteus''. Phylogeny ''Protitanichthys'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny of ''Protitanichthys'': Species ''P. fossatus'' The type species of the genus. The description of ''P. fossatus'' is based on a cranial roof, possibly long, found in the Eifelian-aged D ...
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Panxiosteidae
Panxiosteidae is an extinct family of arthrodire placoderms that lived during the Devonian period. Phylogeny The family Panxiosteidae was erected by Wang in 1979. Members of the family are noted for showing morphologically intermediate traits between coccosteids and dunkleosteids. In the 2010 Carr & Hlavin phylogenetic study, Panxiosteidae was recognized as the sister taxon to the family Dunkleosteidae, which together comprised the superfamily Dunkleosteoidea (one of the three major clades of Eubrachythoraci). The phylogeny of Panxiosteidae from the 2013 Zhu & Zhu study is shown in the cladogram below: However, the subsequent 2016 Zhu ''et al.'' study using a larger morphological dataset recovered Panxiosteidae well outside of Dunkleosteoidea, instead within Coccosteomorphi and then Coccosteoidea as the sister group of Coccosteidae Coccosteidae is a family of arthrodire placoderms from the Early to Late Devonian. Fossils appear in various strata in Europe, North Ame ...
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Jiuchengia
''Jiuchengia longoccipita'' is a coccosteid arthrodire placoderm from the Late Emsian epoch of Wuding, Yunnan. Its skull is similar in form to those of ''Watsonosteus'' and ''Dickosteus ''Dickosteus'' is an extinct genus of coccosteid arthrodire placoderm from the Late Eifelian to Early Givetian stages of the Middle Devonian period. Fossils are found in the Orkneys and Caithness, Scotland. Phylogeny ''Dickosteus'' is a memb ...'', though ''J. longoccipita'' can be easily distinguished from them in that its skull is longer, and has anatomical features in common with homostiids. References Coccosteidae Placoderms of Asia Paleontology in Yunnan Prehistoric animals of China Fossil taxa described in 1983 {{Placoderm-stub ...
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Watsonosteus
''Watsonosteus'' is an extinct genus of coccosteid arthrodire placoderm from the Late Givetian stage of the Middle Devonian period. Fossils are found in the Orkney Islands, Scotland. Phylogeny ''Watsonosteus'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ... of ''Watsonosteus'': References {{Taxonbar, from=Q112327223, from2=Q97371185 Coccosteidae Givetian life Fossils of Scotland Animals described in 1932 ...
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Millerosteus Minor
''Millerosteus'' is an extinct genus of coccosteid arthrodire placoderm from the Early Givetian stage of the Middle Devonian period. Fossils are found in the Orkneys and Caithness, Scotland. Phylogeny ''Millerosteus'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ... of ''Millerosteus'': References {{Taxonbar, from=Q112514447, from2=Q97371187 Coccosteidae Givetian life Fossils of Scotland Fish described in 1959 ...
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Millerosteus
''Millerosteus'' is an extinct genus of coccosteid arthrodire placoderm from the Early Givetian stage of the Middle Devonian period. Fossils are found in the Orkneys and Caithness, Scotland. Phylogeny ''Millerosteus'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ... of ''Millerosteus'': References {{Taxonbar, from=Q112514447, from2=Q97371187 Coccosteidae Givetian life Fossils of Scotland Fish described in 1959 ...
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Dickosteus
''Dickosteus'' is an extinct genus of coccosteid arthrodire placoderm from the Late Eifelian to Early Givetian stages of the Middle Devonian period. Fossils are found in the Orkneys and Caithness, Scotland. Phylogeny ''Dickosteus'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ... of ''Dickosteus'': References {{Taxonbar, from=Q112327258, from2=Q55762836 Coccosteidae Givetian life Fossils of Scotland Fish described in 1963 ...
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Dickosteus Threiplandi
''Dickosteus'' is an extinct genus of coccosteid arthrodire placoderm from the Late Eifelian to Early Givetian stages of the Middle Devonian period. Fossils are found in the Orkneys and Caithness, Scotland. Phylogeny ''Dickosteus'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ... of ''Dickosteus'': References {{Taxonbar, from=Q112327258, from2=Q55762836 Coccosteidae Givetian life Fossils of Scotland Fish described in 1963 ...
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Watsonosteus Fletti
''Watsonosteus'' is an extinct genus of coccosteid arthrodire placoderm from the Late Givetian stage of the Middle Devonian period. Fossils are found in the Orkney Islands, Scotland. Phylogeny ''Watsonosteus'' is a member of the family Coccosteidae, which belongs to the clade Coccosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ... of ''Watsonosteus'': References {{Taxonbar, from=Q112327223, from2=Q97371185 Coccosteidae Givetian life Fossils of Scotland Animals described in 1932 ...
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Coccosteus Cuspidatus
''Coccosteus'' (from el, κόκκος , 'berry' and el, ὀστέον 'bone') is an extinct genus of arthrodire placoderm from the Devonian period. Its fossils have been found throughout Europe and North America. The majority of these have been found in freshwater sediments, though such a large range suggests that they may have been able to enter saltwater. The largest specimens were about , although the average length was . Description Like all other arthrodires, ''Coccosteus'' had a joint between the armor of the body and skull. It also had an internal joint between its neck vertebrae and the back of the skull, allowing for the mouth to be opened even wider. Along with the longer jaws, this allowed ''Coccosteus'' to feed on fairly large prey. The up-and-down movement of the skull also allowed for more water to be pumped through the gills. Possibly, the creature supplemented its diet with organic material filtered from mud using the gills. As with all other arthrodires, ''Cocc ...
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Sister Clade
In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and taxon B are sister groups to each other. Taxa A and B, together with any other extant or extinct descendants of their most recent common ancestor (MRCA), form a monophyletic group, the clade AB. Clade AB and taxon C are also sister groups. Taxa A, B, and C, together with all other descendants of their MRCA form the clade ABC. The whole clade ABC is itself a subtree of a larger tree which offers yet more sister group relationships, both among the leaves and among larger, more deeply rooted clades. The tree structure shown connects through its root to the rest of the universal tree of life. In cladistic standards, taxa A, B, and C may represent specimens, species, genera, or any other taxonomic units. If A and B are at the same taxonomi ...
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