Branchiosauridae
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Branchiosauridae
Branchiosauridae is an extinct family of small amphibamiform temnospondyls with external gills and an overall juvenile appearance. The family has been characterized by hundreds of well-preserved specimens from the Permo-Carboniferous of Middle Europe.Schoch, R.R. 2008. The intrarelationships and evolutionary history of the temnospondyl family branchiosauridae. Journal of Systematic Palaeontology. 6(4):409-431. Specimens represent well defined ontogenetic stages and thus the taxon has been described to display paedomorphy (perennibranchiate). However, more recent work has revealed branchiosaurid taxa that display metamorphosing trajectories.Schoch, R.R. and Frobisch, N.B. 2006. Alternative Pathways in an Extinct Amphibian Clade. Evolution. 60(7):1467-1475 The name Branchiosauridae (“Branchio” in Ancient Greek denoting gills and “saurus” meaning lizard) refers to the retention of gills. Geological/paleoenvironmental information Branchiosaurids mostly inhabited Permo-Carbo ...
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Branchiosaurus Salamandroides
''Branchiosaurus'' (from el, βράγχιον , 'gill' and el, σαῦρος , 'lizard') is a genus of small, lightly built early prehistoric amphibians. Fossils have been discovered in strata dating from the late Pennsylvanian Epoch to the Permian Period. The taxa may be invalid; the material referred to the genus may be juvenile specimens of larger amphibians. This tiny amphibian was very similar to the Rachitomi, differing primarily in size. Other distinguishing characteristics include a cartilaginous, less ossified skeleton and a shorter skull. Clear traces of gills are present in many fossilized samples, hence the name. Originally thought to have vertebrae distinct from rachitomous vertebrae, it was placed in a separate order named Phyllospondyli ("leaf vertebrae"). Later analysis of growth stages showed increasing ossification in larger specimens, which showed that at least some of the species was the larval stage of much larger rachitomes like ''Eryops'', while o ...
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Paleozoic
The Paleozoic (or Palaeozoic) Era is the earliest of three geologic eras of the Phanerozoic Eon. The name ''Paleozoic'' ( ;) was coined by the British geologist Adam Sedgwick in 1838 by combining the Greek words ''palaiós'' (, "old") and ''zōḗ'' (), "life", meaning "ancient life" ). It is the longest of the Phanerozoic eras, lasting from , and is subdivided into six geologic periods (from oldest to youngest): # Cambrian # Ordovician # Silurian # Devonian # Carboniferous # Permian The Paleozoic comes after the Neoproterozoic Era of the Proterozoic Eon and is followed by the Mesozoic Era. The Paleozoic was a time of dramatic geological, climatic, and evolutionary change. The Cambrian witnessed the most rapid and widespread diversification of life in Earth's history, known as the Cambrian explosion, in which most modern phyla first appeared. Arthropods, molluscs, fish, amphibians, reptiles, and synapsids all evolved during the Paleozoic. Life began in the ocean ...
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Plesiomorphic
In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorphy, apomorphy, and synapomorphy, all mean a trait shared between species because they share an ancestral species. Apomorphic and synapomorphic characteristics convey much information about evolutionary clades and can be used to define taxa. However, plesiomorphic and symplesiomorphic characteristics cannot. The term ''symplesiomorphy'' was introduced in 1950 by German entomologist Willi Hennig. Examples A backbone is a plesiomorphic trait shared by birds and mammals, and does not help in placing an animal in one or the other of these two clades. Birds and mammals share this trait because both clades are descended from the same far distant ancestor. Other clades, e.g. snakes, lizards, turtles, fish, frogs, all have backbones and none are either birds no ...
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Synapomorphies
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis T ...
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Branchiosaurus Amblystomus Credner
''Branchiosaurus'' (from el, βράγχιον , 'gill' and el, σαῦρος , 'lizard') is a genus of small, lightly built early prehistoric amphibians. Fossils have been discovered in strata dating from the late Pennsylvanian Epoch to the Permian Period. The taxa may be invalid; the material referred to the genus may be juvenile specimens of larger amphibians. This tiny amphibian was very similar to the Rachitomi, differing primarily in size. Other distinguishing characteristics include a cartilaginous, less ossified skeleton and a shorter skull. Clear traces of gills are present in many fossilized samples, hence the name. Originally thought to have vertebrae distinct from rachitomous vertebrae, it was placed in a separate order named Phyllospondyli ("leaf vertebrae"). Later analysis of growth stages showed increasing ossification in larger specimens, which showed that at least some of the species was the larval stage of much larger rachitomes like ''Eryops'', while o ...
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Monophyletic Group
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species ( extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organis ...
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Paedomorphic
Neoteny (), also called juvenilization,Montagu, A. (1989). Growing Young. Bergin & Garvey: CT. is the delaying or slowing of the physiological, or somatic, development of an organism, typically an animal. Neoteny is found in modern humans compared to other primates. In progenesis or paedogenesis, sexual development is accelerated. Both neoteny and progenesis result in paedomorphism (as having the form typical of children) or paedomorphosis (changing towards forms typical of children), a type of heterochrony. It is the retention in adults of traits previously seen only in the young. Such retention is important in evolutionary biology, domestication and evolutionary developmental biology. Some authors define paedomorphism as the retention of larval traits, as seen in salamanders.Schell, S. C. ''Handbook of Trematodes of North America North of Mexico'', 1985, pg. 22 History and etymology The origins of the concept of neoteny have been traced to the Bible (as argued by Ashley Monta ...
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Larval
A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The larva's appearance is generally very different from the adult form (''e.g.'' caterpillars and butterflies) including different unique structures and organs that do not occur in the adult form. Their diet may also be considerably different. Larvae are frequently adapted to different environments than adults. For example, some larvae such as tadpoles live almost exclusively in aquatic environments, but can live outside water as adult frogs. By living in a distinct environment, larvae may be given shelter from predators and reduce competition for resources with the adult population. Animals in the larval stage will consume food to fuel their transition into the adult form. In some organisms like polychaetes and barnacles, adults are immobile ...
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Gene Flow
In population genetics, gene flow (also known as gene migration or geneflow and allele flow) is the transfer of genetic material from one population to another. If the rate of gene flow is high enough, then two populations will have equivalent allele frequencies and therefore can be considered a single effective population. It has been shown that it takes only "one migrant per generation" to prevent populations from diverging due to drift. Populations can diverge due to selection even when they are exchanging alleles, if the selection pressure is strong enough. Gene flow is an important mechanism for transferring genetic diversity among populations. Migrants change the distribution of genetic diversity among populations, by modifying allele frequencies (the proportion of members carrying a particular variant of a gene). High rates of gene flow can reduce the genetic differentiation between the two groups, increasing homogeneity. For this reason, gene flow has been thought to con ...
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Guadalupian
The Guadalupian is the second and middle series/epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson's Extinction and a significant mass extinction called the end-Capitanian extinction event. The Guadalupian was previously known as the Middle Permian. Name and background The Guadalupian is the second and middle series or epoch of the Permian. Previously called Middle Permian, the name of this epoch is part of a revision of Permian stratigraphy for standard global correlation. The name "Guadalupian" was first proposed in the early 1900s, and approved by the International Subcommission on Permian Stratigraphy in 1996. References to the Middle Permian still exist. The Guadalupian was preceded by the Cisuralian and followed by the Lop ...
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Kaş Formation
Kaş (; el, Αντίφελλος, translit=Antífellos, translit-std=ISO) is a small fishing, diving, yachting and tourist town, and a district of Antalya Province of Turkey, 168 km west of the city of Antalya. As a tourist resort, it is relatively unspoiled. Geography As the tenth largest district of Antalya as of 2021, Kaş is on a hill running down to the Turquoise Coast of southwestern Turkey. The district has a typical Mediterranean climate of hot, dry summers and mild, wet winters, which allows the growth of oranges, lemons and bananas. The lowland areas are also planted with cut flowers and a variety of fruits and vegetables. Many are grown all year round under glass. The hillsides produce honey and almonds, while at high altitudes there are extensive pine forests. The weather is drier at high altitudes. Although agriculture is still important, tourism is the main source of income in the district, which has many hotels and guest houses. About offshore from Kaş i ...
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Wordian
In the geologic timescale, the Wordian is an age or stage of the Permian. It is the middle of three subdivisions of the Guadalupian Epoch or Series. The Wordian lasted between and million years ago (Ma). It was preceded by the Roadian and followed by the Capitanian. Stratigraphy The Wordian Stage was introduced into scientific literature by Johan August Udden in 1916 and was named after the Word Formation of the North American Permian Basin. The Wordian was first used as a stratigraphic subdivision of the Guadalupian in 1961, when both names were still only used regionally in the southern US. The stage was added to the internationally used ISC timescale in 2001.; 1999: ''Proposal of Guadalupian and Component Roadian, Wordian and Capitanian Stages as International Standards for the Middle Permian Series'', Permophiles 34: pp 3–11. The base of the Wordian Stage is defined as the place in the stratigraphic record where fossils of conodont species ''Jinogondolella aserrata'' fi ...
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