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Biological Soil Crust
Biological soil crusts are communities of living organisms on the soil surface in arid and semi-arid ecosystems. They are found throughout the world with varying species composition and cover depending on topography, soil characteristics, climate, plant community, microhabitats, and disturbance regimes. Biological soil crusts perform important ecological roles including carbon fixation, nitrogen fixation and soil stabilization; they alter soil albedo and water relations and affect germination and nutrient levels in vascular plants. They can be damaged by fire, recreational activity, grazing and other disturbances and can require long time periods to recover composition and function. Biological soil crusts are also known as ''biocrusts'' or as ''cryptogamic'', ''microbiotic'', ''microphytic'', or ''cryptobiotic'' soils. Natural history Biology and composition Biological soil crusts are most often composed of fungi, lichens, cyanobacteria, bryophytes, and algae in varying proportions ...
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Hovenweep National Monument
Hovenweep National Monument is located on land in southwestern Colorado and southeastern Utah, between Cortez, Colorado and Blanding, Utah on the Cajon Mesa of the Great Sage Plain. Shallow tributaries run through the wide and deep canyons into the San Juan River.''Hovenweep Visitor Guide''
National Park Service. Retrieved 9-20-2011.
Although Hovenweep National Monument is largely known for the six groups of Ancestral Puebloan villages, there is evidence of occupation by hunter-gatherers from 8,000 to 6,000 B.C. until about AD 200. Later, a succession of early puebloan cultures settled in the area and ...
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Ammonia
Ammonia is an inorganic compound of nitrogen and hydrogen with the formula . A stable binary hydride, and the simplest pnictogen hydride, ammonia is a colourless gas with a distinct pungent smell. Biologically, it is a common nitrogenous waste, particularly among aquatic organisms, and it contributes significantly to the nutritional needs of terrestrial organisms by serving as a precursor to 45% of the world's food and fertilizers. Around 70% of ammonia is used to make fertilisers in various forms and composition, such as urea and Diammonium phosphate. Ammonia in pure form is also applied directly into the soil. Ammonia, either directly or indirectly, is also a building block for the synthesis of many pharmaceutical products and is used in many commercial cleaning products. It is mainly collected by downward displacement of both air and water. Although common in nature—both terrestrially and in the outer planets of the Solar System—and in wide use, ammonia i ...
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Cryptobiotic Soil Crust In Natural Bridges National Monument Near Sipapu 20100906 - Number 5
Cryptobiosis or anabiosis is a metabolic state of life entered by an organism in response to adverse environmental conditions such as desiccation, freezing, and oxygen deficiency. In the cryptobiotic state, all measurable metabolic processes stop, preventing reproduction, development, and repair. When environmental conditions return to being hospitable, the organism will return to its metabolic state of life as it was prior to the cryptobiosis. Forms Anhydrobiosis Anhydrobiosis is the most studied form of cryptobiosis and occurs in situations of extreme desiccation. The term ''anhydrobiosis'' derives from the Greek for "life without water" and is most commonly used for the desiccation tolerance observed in certain invertebrate animals such as bdelloid rotifers, tardigrades, brine shrimp, nematodes, and at least one insect, a species of chironomid (''Polypedilum vanderplanki''). However, other life forms exhibit desiccation tolerance. These include the resurrection plant ' ...
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Vascular Plants
Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They also have a specialized non-lignified tissue (the phloem) to conduct products of photosynthesis. Vascular plants include the clubmosses, horsetails, ferns, gymnosperms (including conifers), and angiosperms (flowering plants). Scientific names for the group include Tracheophyta, Tracheobionta and Equisetopsida ''sensu lato''. Some early land plants (the rhyniophytes) had less developed vascular tissue; the term eutracheophyte has been used for all other vascular plants, including all living ones. Historically, vascular plants were known as "higher plants", as it was believed that they were further evolved than other plants due to being more complex organisms. However, this is an antiquated remnant of the obsolete scala naturae, and the ...
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Green Algae
The green algae (singular: green alga) are a group consisting of the Prasinodermophyta and its unnamed sister which contains the Chlorophyta and Charophyta/ Streptophyta. The land plants ( Embryophytes) have emerged deep in the Charophyte alga as sister of the Zygnematophyceae. Since the realization that the Embryophytes emerged within the green algae, some authors are starting to properly include them. The completed clade that includes both green algae and embryophytes is monophyletic and is referred to as the clade Viridiplantae and as the kingdom Plantae. The green algae include unicellular and colonial flagellates, most with two flagella per cell, as well as various colonial, coccoid and filamentous forms, and macroscopic, multicellular seaweeds. There are about 22,000 species of green algae. Many species live most of their lives as single cells, while other species form coenobia (colonies), long filaments, or highly differentiated macroscopic seaweeds. A few oth ...
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Hyphae
A hypha (; ) is a long, branching, filamentous structure of a fungus, oomycete, or actinobacterium. In most fungi, hyphae are the main mode of vegetative growth, and are collectively called a mycelium. Structure A hypha consists of one or more cells surrounded by a tubular cell wall. In most fungi, hyphae are divided into cells by internal cross-walls called "septa" (singular septum). Septa are usually perforated by pores large enough for ribosomes, mitochondria, and sometimes nuclei to flow between cells. The major structural polymer in fungal cell walls is typically chitin, in contrast to plants and oomycetes that have cellulosic cell walls. Some fungi have aseptate hyphae, meaning their hyphae are not partitioned by septa. Hyphae have an average diameter of 4–6 µm. Growth Hyphae grow at their tips. During tip growth, cell walls are extended by the external assembly and polymerization of cell wall components, and the internal production of new cell membrane. Th ...
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Black Yeast
“Black yeasts”, sometimes also black fungi, dematiaceous fungi, microcolonial fungi or meristematic fungi is a diverse group of slow-growing microfungi which reproduce mostly asexually ( fungi imperfecti). Only few genera reproduce by budding cells, while in others hyphal or meristematic (isodiametric) reproduction is preponderant. Black yeasts share some distinctive characteristics, in particular a dark colouration ( melanisation) of their cell wall. Morphological plasticity, incrustation of the cell wall with melanins and presence of other protective substances like carotenoids and mycosporines represent passive physiological adaptations which enable black fungi to be highly resistant against environmental stresses. The term " polyextremotolerance" has been introduced to describe this phenotype, a good example of which is the species '' Aureobasidium pullulans''. Presence of 1,8-dihydroxynaphthalene melanin in the cell wall confers to the microfungi their characteristic ol ...
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Melanin
Melanin (; from el, μέλας, melas, black, dark) is a broad term for a group of natural pigments found in most organisms. Eumelanin is produced through a multistage chemical process known as melanogenesis, where the oxidation of the amino acid tyrosine is followed by polymerization. The melanin pigments are produced in a specialized group of cells known as melanocytes. Functionally, eumelanin serves as protection against UV radiation. There are five basic types of melanin: eumelanin, pheomelanin, neuromelanin, allomelanin and pyomelanin. The most common type is eumelanin, of which there are two types— brown eumelanin and black eumelanin. Pheomelanin, which is produced when melanocytes are malfunctioning due to derivation of the gene to its recessive format is a cysteine-derivative that contains poly benzothiazine portions that are largely responsible for the of red yellow tint given to some skin or hair colors. Neuromelanin is found in the brain. Research has been ...
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Symbiosis
Symbiosis (from Greek , , "living together", from , , "together", and , bíōsis, "living") is any type of a close and long-term biological interaction between two different biological organisms, be it mutualistic, commensalistic, or parasitic. The organisms, each termed a symbiont, must be of different species. In 1879, Heinrich Anton de Bary defined it as "the living together of unlike organisms". The term was subject to a century-long debate about whether it should specifically denote mutualism, as in lichens. Biologists have now abandoned that restriction. Symbiosis can be obligatory, which means that one or more of the symbionts depend on each other for survival, or facultative (optional), when they can generally live independently. Symbiosis is also classified by physical attachment. When symbionts form a single body it is called conjunctive symbiosis, while all other arrangements are called disjunctive symbiosis."symbiosis." Dorland's Illustrated Medical Dictionary ...
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Microfungi
Microfungi or micromycetes are fungi—eukaryotic organisms such as molds, mildews and rusts—which have microscopic spore-producing structures. They exhibit tube tip-growth and have cell walls composed of chitin, a polymer of ''N''-acetylglucosamine. Microfungi are a paraphyletic group, distinguished from macrofungi only by the absence of a large, multicellular fruiting body. They are ubiquitous in all terrestrial and freshwater and marine environments, and grow in plants, soil, water, insects, cattle rumens, hair, and skin. Most of the fungal body consists of microscopic threads, called hyphae, extending through the substrate in which it grows. The mycelia of microfungi produce spores that are carried by the air, spreading the fungus. Many microfungi species are benign, existing as soil saprotrophs, for example, largely unobserved by humans. Many thousands of microfungal species occur in lichens, forming symbiotic relationships with algae. Other microfungi, such as thos ...
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Lichens And Nitrogen Cycling
Some types of lichen are able to fix nitrogen from the atmosphere. This process relies on the presence of cyanobacteria as a partner species within the lichen. The ability to fix nitrogen enables lichen to live in nutrient-poor environments. Lichen can also extract nitrogen from the rocks on which they grow. Nitrogen fixation, and hence the abundance of lichen and their host plants, may be decreased by application of nitrogen-based agricultural fertilizer and by atmospheric pollution. The nitrogen cycle The nitrogen cycle is one of the Earth's biogeochemical cycles. It involves the conversion of nitrogen into different chemical forms. The main processes of the nitrogen cycle are the fixation, ammonification, nitrification, and denitrification. As one of the macronutrients, nitrogen plays an important role in plant growth. The nitrogen cycle is affected by environmental factors. For example, in the subarctic heath, increase in temperature can cause nitrogen fixation to increase or ...
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Squamulose
A squamulose lichen is a lichen that is composed of small, often overlapping "scales" called squamules. If they are raised from the substrate and appear leafy, the lichen may appear to be a foliose lichen, but the underside does not have a "skin" (cortex), as foliose lichens do. Squamulose lichens are composed of flattish units that are usually tightly clustered. They are like an intermediate between crustose and foliose lichens. Examples of squamulose lichen include ''Vahliella leucophaea'', ''Cladonia subcervicornis'' and ''Lichenomphalia hudsoniana ''Lichenomphalia hudsoniana'' is a species of basidiolichen in the family Hygrophoraceae. It is widely distributed in alpine and arctic regions of the world, where it grows on moist soil amongst moss. Taxonomy The species was first species descr ...''. References Lichenology {{lichen-stub ...
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