Anteosaurid
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Anteosaurid
Anteosauridae is an extinct family of large carnivorous dinocephalian therapsids that are known from the Middle Permian of Asia, Africa, and South America.These animals were by far the largest predators of the Permian period, with skulls reaching 80 cm in length in adult individuals, far larger than the biggest gorgonopsian. Description Anteosaurids are characterized by very large pointed incisors and canines, bulbous spatulate (spoon shaped) postcanines, a very strongly upturned margin of the premaxilla, so the front of mouth curves strongly upwards, and a long, very robust lower jaw (Boonstra 1963). Anteosaurids distinguished from ''Brithopus'' and other non-anteosaurid anteosaurians by the presence of a large thickened region or "boss" on the side of the angle of the lower jaw; this was probably used in intraspecific behaviour. In ''Doliosauriscus'' and '' Anteosaurus'', not only was this boss very prominent, but the bones were very thick and rugose. The same situat ...
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Anteosaurus
''Anteosaurus'' (meaning "Antaeus reptile") is an extinct genus of large carnivorous dinocephalian synapsid. It lived at the end of the Guadalupian (= Middle Permian) during the Capitanian stage, about 265 to 260 million years ago in what is now South Africa. It is mainly known by cranial remains and few postcranial bones. With its skull reaching in length and a body size estimated at more than in length, and in weight, ''Anteosaurus'' was the largest known carnivorous non-mammalian synapsid and the largest terrestrial predator of the Permian period. Occupying the top of the food chain in the Middle Permian, its skull, jaws and teeth show adaptations to capture large prey like the giants titanosuchids and tapinocephalids dinocephalians and large pareiasaurs. As in many other dinocephalians the cranial bones of ''Anteosaurus'' are pachyostosed, but to a lesser extent than in tapinocephalid dinocephalians. In ''Anteosaurus'', pachyostosis mainly occurs in the form of horn-sha ...
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Paranteosaurus
''Anteosaurus'' (meaning "Antaeus reptile") is an extinct genus of large carnivorous dinocephalian synapsid. It lived at the end of the Guadalupian (= Middle Permian) during the Capitanian stage, about 265 to 260 million years ago in what is now South Africa. It is mainly known by cranial remains and few postcranial bones. With its skull reaching in length and a body size estimated at more than in length, and in weight, ''Anteosaurus'' was the largest known carnivorous non-mammalian synapsid and the largest terrestrial predator of the Permian period. Occupying the top of the food chain in the Middle Permian, its skull, jaws and teeth show adaptations to capture large prey like the giants titanosuchids and tapinocephalids dinocephalians and large pareiasaurs. As in many other dinocephalians the cranial bones of ''Anteosaurus'' are pachyostosed, but to a lesser extent than in tapinocephalid dinocephalians. In ''Anteosaurus'', pachyostosis mainly occurs in the form of horn-s ...
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Anteosauria
Anteosaurs are a group of large, primitive carnivorous dinocephalian therapsids with large canines and incisors and short limbs, that are known from the Middle Permian of South Africa, Russia, China, and Brazil. Some grew very large, with skulls long, and were the largest predators of their time. They died out at the end of the Middle Permian, possibly as a result of the extinction of the herbivorous Tapinocephalia on which they may have fed. Description The Anteosauria are distinguished from the Tapinocephalia by a number of features, such as very large canines, cheek teeth with bulbous crowns, and an upturning of the premaxilla, so that the front of mouth curves strongly upwards. There is a tendency especially in more advanced forms such as ''Anteosaurus'' towards thickening of the bones of the top of the skull, indicating head-butting behaviour. There is a large canal for the pineal organ (third eye); probably tied in with the animal's diurnal and seasonal cycles. The should ...
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Anteosaurinae
Anteosaurinae is an extinct subfamily of dinocephalian therapsids. It is one of two subfamilies in the family Anteosauridae, the other being Syodontinae. Description These are very specialized, very large anteosaurs. The postcanine teeth are further reduced. Deepening of the postorbital region of the skull (behind the eyes) produced a larger temporal opening, indicating more muscle mass. The boss on the angular (rear of the jaw) has become very prominent, again, another sign of powerful jaw muscles. These huge animals were clearly formidable predators. In the Anteosaurinae, pachyostosis is taken to extremes. The dorsal (upper) surface of the nasal, frontal, and postfrontals (around and between/above the eyes) is thickened and rugose in the same manner as the tapinocephalids. Nevertheless these animals are too specialized and too late in time to have been the ancestors of the herbivorous tapinocephalids, so these characteristics evolved independently. Boonstra notes that the h ...
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Dinocephalia
Dinocephalians (terrible heads) are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinocephalians included herbivorous, carnivorous, and omnivorous forms. Many species had thickened skulls with many knobs and bony projections. Dinocephalians were the first non-mammalian therapsids to be scientifically described and their fossils are known from Russia, China, Brazil, South Africa, Zimbabwe, and Tanzania. Description Apart from the Biarmosuchians, the dinocephalians are the least advanced therapsids, although still uniquely specialised in their own way. They retain a number of primitive characteristics (e.g. no secondary palate, small dentary) shared with their pelycosaur ancestors, although they are also more advanced in possessing therapsid adaptations like the expansion of the ilium and more erect limbs. They include carni ...
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Syodontinae
Syodontinae is a group of dinocephalian therapsids. It is one of two subfamilies in the family Anteosauridae, the other being Anteosaurinae. They are known from the Middle Permian Period of what is now Russia and South Africa. One of the best known syodontines is ''Syodon'' from Russia. The South African form ''Australosyodon'', is one of the earliest known Gondwanan anteosaurs. Description Syodontines lack the boss on the lower jaw that characterises the related anteosaurines, and they have often been considered more primitive in this respect. Classification Below is a cladogram showing syodontine relationships from a 2012 phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ... study of anteosaurians: References Anteosaurs Guadalupian first appearances Guada ...
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Dinocephalia
Dinocephalians (terrible heads) are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinocephalians included herbivorous, carnivorous, and omnivorous forms. Many species had thickened skulls with many knobs and bony projections. Dinocephalians were the first non-mammalian therapsids to be scientifically described and their fossils are known from Russia, China, Brazil, South Africa, Zimbabwe, and Tanzania. Description Apart from the Biarmosuchians, the dinocephalians are the least advanced therapsids, although still uniquely specialised in their own way. They retain a number of primitive characteristics (e.g. no secondary palate, small dentary) shared with their pelycosaur ancestors, although they are also more advanced in possessing therapsid adaptations like the expansion of the ilium and more erect limbs. They include carni ...
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Titanophoneus
''Titanophoneus'' ("titanic murderer") is an extinct genus of carnivorous dinocephalian therapsid from the Middle Permian. It is classified within the family Anteosauridae. The type species is ''Titanophoneus potens''. Remains of ''Titanophoneus'' have been found at Isheevo in Russia. Description An adult skull would have reached 80 cm with a heavy long snout. The long tail and short limbs show the species to be a primitive therapsid unlike '' Inostrancevia'' which was more advanced. The structure of the limbs and the density of the bone are designed for a sprawling stance. The temporal opening is more advanced than Estemmenosuchidae but less advanced than ''Inostrancevia''. The teeth are large with 12 large palate incisors followed by 2 canines and various smaller back teeth. The lower palate is the same as the upper but without the canine teeth. The appearance of ''Titanophoneus'' is reminiscent to the sphenacodontid pelycosaurs, which included ''Dimetrodon''. Gallery im ...
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Doliosauriscus
''Titanophoneus'' ("titanic murderer") is an extinct genus of carnivorous dinocephalian therapsid from the Middle Permian. It is classified within the family Anteosauridae. The type species is ''Titanophoneus potens''. Remains of ''Titanophoneus'' have been found at Isheevo in Russia. Description An adult skull would have reached 80 cm with a heavy long snout. The long tail and short limbs show the species to be a primitive therapsid unlike ''Inostrancevia'' which was more advanced. The structure of the limbs and the density of the bone are designed for a sprawling stance. The temporal opening is more advanced than Estemmenosuchidae but less advanced than ''Inostrancevia''. The teeth are large with 12 large palate incisors followed by 2 canines and various smaller back teeth. The lower palate is the same as the upper but without the canine teeth. The appearance of ''Titanophoneus'' is reminiscent to the sphenacodontid pelycosaurs, which included ''Dimetrodon''. Gallery imag ...
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Tapinocephalidae
Tapinocephalidae was an advanced family of tapinocephalians. It is defined as the clade containing ''Ulemosaurus'', '' Tapinocaninus'', and the Tapinocephalinae. They are known from both Russia and South Africa. In all probability, the Tapinocephalidae had a worldwide (Pangean) distribution. They flourished briefly during the Wordian and Capitanian ages, radiating into several lineages, existing simultaneously, and differing mainly in details of the skull and, to an even lesser degree, the skeleton. It is not clear how such similar animals could each find their own ecological niche, but such was obviously the case. There is a parallel here with the hadrosaur and ceratopsian dinosaurs of the Late Cretaceous. The cause of their abrupt extinction is not clear, since other smaller animals, and even the pareiasaurs, were not affected. Quite probably, like the extinction of the late Pleistocene megafauna, a number of factors were involved. Description The body is deep and capacious, al ...
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Ivan Yefremov
Ivan Antonovich (real patronymic Antipovich) Yefremov ( ru , Ива́н Анто́нович (Анти́пович) Ефре́мов; April 23, 1908 – October 5, 1972; last name sometimes transliterated as Efremov) was a Soviet paleontologist, science-fiction author and social thinker. He founded taphonomy, the study of fossilization patterns. Biography He was born in the village of Vyritsa in Saint Petersburg Governorate on April 23, 1908. His parents divorced during the Russian Revolution. His mother married a Red Army commander and left the children in Kherson to be cared for by an aunt who soon died of typhus. Yefremov survived on his own for some time, after which he joined a Red Army unit as a "son of the regiment" and went to Perekop with it. In 1921, he was discharged and went to Petrograd (today's Saint Petersburg) to study. He completed his education there while combining his studies with a variety of odd jobs. He later commented that "the Revolution was als ...
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Brithopodidae
''Brithopus'' is an extinct genus of dinocephalian therapsids. It contains a single species, ''Brithopus priscus'', known from fragmentary remains found in the Copper Sandstones near Isheevo, Russia. Description ''Brithopus'' was fairly large, reaching a length of 2.5–3 m (8–10 ft). The skull was similar to ''Titanophoneus'', but more massive and heavily built.Olson, E.C. (1962). "Late Permian terrestrial vertebrates, U.S.A. and U.S.S.R." ''Transactions of the American Philosophical Society'', New Series, 52: 1–224. Classification ''B. priscus'' was first named in 1838 and was traditionally classified in the Anteosauria, a group of carnivorous dinocephalians. ''Brithopus'' served as the basis for the family Brithopodidae, which once included many anteosaurian species. Because it is based on fragmentary material, ''Brithopus'' is regarded as a ''nomen dubium'' by some researchers. ''Brithopus'' was later considered a possible estemmenosuchid,Kammerer, C. F. 2010. Syste ...
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