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Syodontinae
Syodontinae is a group of dinocephalian therapsids. It is one of two subfamilies in the family Anteosauridae, the other being Anteosaurinae. They are known from the Middle Permian Period of what is now Russia and South Africa. One of the best known syodontines is ''Syodon'' from Russia. The South African form ''Australosyodon'', is one of the earliest known Gondwanan anteosaurs. Description Syodontines lack the boss on the lower jaw that characterises the related anteosaurines, and they have often been considered more primitive in this respect. Classification Below is a cladogram showing syodontine relationships from a 2012 phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ... study of anteosaurians: References Anteosaurs Guadalupian first appearances Guada ...
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Sinophoneus Yumenensis
''Sinophoneus'' is an extinct genus of carnivorous dinocephalian therapsid belonging to the family Anteosauridae. It lived 272 to 270 million years ago at the beginning of the Middle Permian (Lower Roadian) in what is now the Gansu Province in northern China. It is known by a skull of an adult individual (the holotype GMV1601), as well as by many skulls of juvenile specimens. The latter were first considered as belonging to a different animal, named ''Stenocybus'', before being reinterpreted as immature ''Sinophoneus''. ''Sinophoneus'' shows a combination of characters present in other anteosaurs. Its bulbous profile snout and external nostrils located in front of the canine are reminiscent of the basal anteosaur ''Archaeosyodon'', while its massive transerse pterygoids processes with enlarged distal ends are more similar to the more derived anteosaurs ''Anteosaurus'' and ''Titanophoneus''. First phylogenetic analyzes identified ''Sinophoneus'' as the most basal Anteosaurinae. A ...
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Anteosaurus Magnificus
''Anteosaurus'' (meaning " Antaeus reptile") is an extinct genus of large carnivorous dinocephalian synapsid. It lived at the end of the Guadalupian (= Middle Permian) during the Capitanian stage, about 265 to 260 million years ago in what is now South Africa. It is mainly known by cranial remains and few postcranial bones. With its skull reaching in length and a body size estimated at more than in length, and in weight, ''Anteosaurus'' was the largest known carnivorous non-mammalian synapsid and the largest terrestrial predator of the Permian period. Occupying the top of the food chain in the Middle Permian, its skull, jaws and teeth show adaptations to capture large prey like the giants titanosuchids and tapinocephalids dinocephalians and large pareiasaurs. As in many other dinocephalians the cranial bones of ''Anteosaurus'' are pachyostosed, but to a lesser extent than in tapinocephalid dinocephalians. In ''Anteosaurus'', pachyostosis mainly occurs in the form of horn- ...
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Pampaphoneus
''Pampaphoneus'' is an extinct genus of carnivorous dinocephalian therapsid belonging to the family Anteosauridae. It lived 268 to 265 million years ago during the Wordian age of the Guadalupian (= middle Permian) period in what is now Brazil. ''Pampaphoneus'' is known by an almost complete skull with the lower jaw still articulated, discovered on the lands of the Boqueirão Farm, near the city of São Gabriel, in the state of Rio Grande do Sul. A second specimen from the same locality was reported in 2019 and 2020 but has not yet been described. It is composed of a skull associated with postcranial remains. It is the first South American species of dinocephalian to have been described. The group was previously known in South America only by a few isolated teeth and a jaw fragment reported in 2000 in the same region of Brazil. Phylogenetic analysis conducted by Cisneros and colleagues reveals that ''Pampaphoneus'' is closely related to anteosaurs from European Russia, indicating ...
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Pampaphoneus Biccai
''Pampaphoneus'' is an extinct genus of carnivorous dinocephalian therapsid belonging to the family Anteosauridae. It lived 268 to 265 million years ago during the Wordian age of the Guadalupian (= middle Permian) period in what is now Brazil. ''Pampaphoneus'' is known by an almost complete skull with the lower jaw still articulated, discovered on the lands of the Boqueirão Farm, near the city of São Gabriel, in the state of Rio Grande do Sul. A second specimen from the same locality was reported in 2019 and 2020 but has not yet been described. It is composed of a skull associated with postcranial remains. It is the first South American species of dinocephalian to have been described. The group was previously known in South America only by a few isolated teeth and a jaw fragment reported in 2000 in the same region of Brazil. Phylogenetic analysis conducted by Cisneros and colleagues reveals that ''Pampaphoneus'' is closely related to anteosaurs from European Russia, indicating ...
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Anteosauridae
Anteosauridae is an extinct family of large carnivorous dinocephalian therapsids that are known from the Middle Permian of Asia, Africa, and South America.These animals were by far the largest predators of the Permian period, with skulls reaching 80 cm in length in adult individuals, far larger than the biggest gorgonopsian. Description Anteosaurids are characterized by very large pointed incisors and canines, bulbous spatulate (spoon shaped) postcanines, a very strongly upturned margin of the premaxilla, so the front of mouth curves strongly upwards, and a long, very robust lower jaw (Boonstra 1963). Anteosaurids distinguished from ''Brithopus'' and other non-anteosaurid anteosaurians by the presence of a large thickened region or "boss" on the side of the angle of the lower jaw; this was probably used in intraspecific behaviour. In '' Doliosauriscus'' and ''Anteosaurus'', not only was this boss very prominent, but the bones were very thick and rugose. The same situation ...
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Australosyodon
''Australosyodon'' is an extinct genus of dinocephalian therapsids from the middle Permian of South Africa. The first fossil was discovered in the 1980s near the village of Prince Albert Road in the Karoo region of South Africa. The genus is closely related to the genus ''Syodon'', found in the Russian Ischejewo fauna, and is thus regarded as the earliest example of an anteosaurid Dinocephalia outside of Russia. ''Australosyodon'' fossils have been recovered from the Eodicynodon Assemblage Zone, the lowest bank of the South African Beaufort Group, indicating the presence of primitive early therapsids in the southern hemisphere. The genus was first described by Bruce S. Rubidge in 1994. Description ''Australosyodon'' was a medium-sized dinocephalian. The skull was high and narrow, with a length of 26 cm, indicating a total body length of approximately 1.8 m (6 ft). The top of the skull displayed the thickening typical of dinocephalians. Overall there are many ...
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Anteosaurs
Anteosaurs are a group of large, primitive carnivorous dinocephalian therapsids with large canines and incisors and short limbs, that are known from the Middle Permian of South Africa, Russia, China, and Brazil. Some grew very large, with skulls long, and were the largest predators of their time. They died out at the end of the Middle Permian, possibly as a result of the extinction of the herbivorous Tapinocephalia on which they may have fed. Description The Anteosauria are distinguished from the Tapinocephalia by a number of features, such as very large canines, cheek teeth with bulbous crowns, and an upturning of the premaxilla, so that the front of mouth curves strongly upwards. There is a tendency especially in more advanced forms such as ''Anteosaurus'' towards thickening of the bones of the top of the skull, indicating head-butting behaviour. There is a large canal for the pineal organ (third eye); probably tied in with the animal's diurnal and seasonal cycles. The shoul ...
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Australosyodon Nyaphuli
''Australosyodon'' is an extinct genus of dinocephalian therapsids from the middle Permian of South Africa. The first fossil was discovered in the 1980s near the village of Prince Albert Road in the Karoo region of South Africa. The genus is closely related to the genus ''Syodon'', found in the Russian Ischejewo fauna, and is thus regarded as the earliest example of an anteosaurid Dinocephalia outside of Russia. ''Australosyodon'' fossils have been recovered from the Eodicynodon Assemblage Zone, the lowest bank of the South African Beaufort Group, indicating the presence of primitive early therapsids in the southern hemisphere. The genus was first described by Bruce S. Rubidge in 1994. Description ''Australosyodon'' was a medium-sized dinocephalian. The skull was high and narrow, with a length of 26 cm, indicating a total body length of approximately 1.8 m (6 ft). The top of the skull displayed the thickening typical of dinocephalians. Overall there are many ...
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Anteosaurinae
Anteosaurinae is an extinct subfamily of dinocephalian therapsids. It is one of two subfamilies in the family Anteosauridae, the other being Syodontinae. Description These are very specialized, very large anteosaurs. The postcanine teeth are further reduced. Deepening of the postorbital region of the skull (behind the eyes) produced a larger temporal opening, indicating more muscle mass. The boss on the angular (rear of the jaw) has become very prominent, again, another sign of powerful jaw muscles. These huge animals were clearly formidable predators. In the Anteosaurinae, pachyostosis is taken to extremes. The dorsal (upper) surface of the nasal, frontal, and postfrontals (around and between/above the eyes) is thickened and rugose in the same manner as the tapinocephalids. Nevertheless these animals are too specialized and too late in time to have been the ancestors of the herbivorous tapinocephalids, so these characteristics evolved independently. Boonstra notes that the h ...
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Middle Permian
The Guadalupian is the second and middle series/epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson's Extinction and a significant mass extinction called the end-Capitanian extinction event. The Guadalupian was previously known as the Middle Permian. Name and background The Guadalupian is the second and middle series or epoch of the Permian. Previously called Middle Permian, the name of this epoch is part of a revision of Permian stratigraphy for standard global correlation. The name "Guadalupian" was first proposed in the early 1900s, and approved by the International Subcommission on Permian Stratigraphy in 1996. References to the Middle Permian still exist. The Guadalupian was preceded by the Cisuralian and followed by the Lop ...
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Titanophoneus Adamanteus
''Titanophoneus'' ("titanic murderer") is an extinct genus of carnivorous dinocephalian therapsid from the Middle Permian. It is classified within the family Anteosauridae. The type species is ''Titanophoneus potens''. Remains of ''Titanophoneus'' have been found at Isheevo in Russia. Description An adult skull would have reached 80 cm with a heavy long snout. The long tail and short limbs show the species to be a primitive therapsid unlike ''Inostrancevia'' which was more advanced. The structure of the limbs and the density of the bone are designed for a sprawling stance. The temporal opening is more advanced than Estemmenosuchidae but less advanced than ''Inostrancevia''. The teeth are large with 12 large palate incisors followed by 2 canines and various smaller back teeth. The lower palate is the same as the upper but without the canine teeth. The appearance of ''Titanophoneus'' is reminiscent to the sphenacodontid pelycosaurs, which included ''Dimetrodon''. Gallery ima ...
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Titanophoneus Potens
''Titanophoneus'' ("titanic murderer") is an extinct genus of carnivorous dinocephalian therapsid from the Middle Permian. It is classified within the family Anteosauridae. The type species is ''Titanophoneus potens''. Remains of ''Titanophoneus'' have been found at Isheevo in Russia. Description An adult skull would have reached 80 cm with a heavy long snout. The long tail and short limbs show the species to be a primitive therapsid unlike ''Inostrancevia'' which was more advanced. The structure of the limbs and the density of the bone are designed for a sprawling stance. The temporal opening is more advanced than Estemmenosuchidae but less advanced than ''Inostrancevia''. The teeth are large with 12 large palate incisors followed by 2 canines and various smaller back teeth. The lower palate is the same as the upper but without the canine teeth. The appearance of ''Titanophoneus'' is reminiscent to the sphenacodontid pelycosaurs, which included ''Dimetrodon''. Gallery imag ...
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