Anomocephaloids
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Anomocephaloids
Anomocephaloidea is a clade (evolutionary grouping) of anomodont therapsids that existed in Gondwana during the Middle Permian and includes two species, ''Anomocephalus africanus'' from South Africa and ''Tiarajudens eccentricus'' from Brazil, both of which are characterized by large body size and teeth that fit tightly together or occlude. Anomocephaloidea is among the most basal groups of anomodonts, the other being Venyukovioidea, which differs in being a Laurasian clade of mostly small-bodied species. Anomocephaloidea was named in 2011 with the discovery of ''Tiarajudens''; ''Anomocephalus'' had been known since 1999, but was unique among anomodonts until ''Tiarajudens'' was described. Both ''Anomocephalus'' and ''Tiarajudens'' were herbivores, although the latter possessed a pair of saber-like canine teeth that may have been used in display or combat with other individuals of the same species. Although Anomocephaloidea was short-lived and had a limited geographic distri ...
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Venyukovioidea
Venyukovioidea is an infraorder of anomodont therapsids related to dicynodonts from the Permian of Russia. They have also known as Venjukovioidea, as well as by the similar names Venyukoviamorpha or Venjukoviamorpha in literature. This in part owes to a misspelling by Russian palaeontologist Ivan Efremov in 1940 when he mistakenly spelt ''Venyukovia'', the namesake of the group, with a 'j' instead of a 'y' (i.e. ''Venjukovia''), which permeated through subsequent therapsid literature before the mistake was caught and corrected. The order Ulemicia has also been coined for a similar taxonomic concept in Russian scientific literature, which notably excludes ''Suminia'' and ''Parasuminia''. Venyukovioidea includes the genera ''Venyukovia'', ''Otsheria'', ''Ulemica'', ''Suminia'' and ''Parasuminia'', all from Western Siberia. Historically, some of these genera have been placed in various families and subfamilies, including the Venyukoviidae/Venjukoviidae, Otsheriidae, and Ulemiciidae. ...
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Middle Permian
The Guadalupian is the second and middle series/epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson's Extinction and a significant mass extinction called the end-Capitanian extinction event. The Guadalupian was previously known as the Middle Permian. Name and background The Guadalupian is the second and middle series or epoch of the Permian. Previously called Middle Permian, the name of this epoch is part of a revision of Permian stratigraphy for standard global correlation. The name "Guadalupian" was first proposed in the early 1900s, and approved by the International Subcommission on Permian Stratigraphy in 1996. References to the Middle Permian still exist. The Guadalupian was preceded by the Cisuralian and followed by the Lop ...
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Evolutionary Radiation
An evolutionary radiation is an increase in taxonomic diversity that is caused by elevated rates of speciation, that may or may not be associated with an increase in morphological disparity. Radiations may affect one clade or many, and be rapid or gradual; where they are rapid, and driven by a single lineage's adaptation to their environment, they are termed adaptive radiations. Examples Perhaps the most familiar example of an evolutionary radiation is that of placental mammals immediately after the extinction of the non-avian dinosaurs at the end of the Cretaceous, about 66 million years ago. At that time, the placental mammals were mostly small, insect-eating animals similar in size and shape to modern shrews. By the Eocene (58–37 million years ago), they had evolved into such diverse forms as bats, whales, and horses. Other familiar radiations include the Avalon Explosion, the Cambrian Explosion, the Great Ordovician Biodiversification Event, the Carboniferous-Earliest Perm ...
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Titanophoneus
''Titanophoneus'' ("titanic murderer") is an extinct genus of carnivorous dinocephalian therapsid from the Middle Permian. It is classified within the family Anteosauridae. The type species is ''Titanophoneus potens''. Remains of ''Titanophoneus'' have been found at Isheevo in Russia. Description An adult skull would have reached 80 cm with a heavy long snout. The long tail and short limbs show the species to be a primitive therapsid unlike '' Inostrancevia'' which was more advanced. The structure of the limbs and the density of the bone are designed for a sprawling stance. The temporal opening is more advanced than Estemmenosuchidae but less advanced than ''Inostrancevia''. The teeth are large with 12 large palate incisors followed by 2 canines and various smaller back teeth. The lower palate is the same as the upper but without the canine teeth. The appearance of ''Titanophoneus'' is reminiscent to the sphenacodontid pelycosaurs, which included ''Dimetrodon''. Gallery im ...
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Tapinocaninus
''Tapinocaninus'' is an extinct genus of therapsids in the family Tapinocephalidae, of which it is the most basal member. Only one species is known, ''Tapinocaninus pamelae'' (meaning "Pam's humble canine"). Fossils have been found dating from the Middle Permian (Wordian age). Description This species is known from several skulls. It was a large animal, reaching up to in body mass. It is assumed to be either a herbivore or an omnivore. This dinocephalian has a heterodont dentition, consisting of incisors, canines and postcanines. Its canine teeth lack a definite crushing heel. The maxillary and premaxillary dentition is only partially preserved. The species is named in honour of Rubidge's mother. Discovery Fossils have been found in the Eodicynodon Assemblage Zone The ''Eodicynodon'' Assemblage Zone is a tetrapod assemblage zone or biozone which correlates to the Abrahamskraal Formation, Adelaide Subgroup of the Beaufort Group, a fossiliferous and geologically important geo ...
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Estemmenosuchus
''Estemmenosuchus'' (meaning "crowned crocodile" in Greek) is an extinct genus of large, early omnivorous therapsid. It is believed and interpreted to have lived during the middle part of the Middle Permian around 267 million years ago. The two species, ''E. uralensis'' and ''E. mirabilis'', are characterised by distinctive horn-like structures, which were probably used for intra-specific display. Both species of ''Estemmenosuchus'' are from the Perm (or Cis-Urals) region of Russia. Two other estemmenosuchids, ''Anoplosuchus'' and ''Zopherosuchus'', are now considered females of the species ''E. uralensis''. There were many complete and incomplete skeletons found together. Description ''Estemmenosuchus'' could reach a body length of more than . Its skull was long and massive, up to in length, and possessed several sets of large horns, somewhat similar to the antlers of a moose, growing upward and outward from the sides and top of the head. The animal had a sprawling posture a ...
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Herpetoskylax
''Herpetoskylax'' is an extinct genus of biarmosuchians which existed in South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring coun .... The type species is ''Herpetoskylax hopsoni''. It lived in the Late Permian Period. The genus name means ‘reptile-puppy’, from the Ancient Greek ' (, ‘creeping animal’) and ' (, ‘young dog’). The juxtaposition of reptilian and mammalian names highlights the transitional characters of therapsids. The type specimen is CGP 1/67, a skull. References Biarmosuchians Prehistoric therapsid genera Lopingian synapsids of Africa Fossil taxa described in 2006 Taxa named by Christian Sidor Lopingian genus first appearances Lopingian genus extinctions {{Paleo-therapsid-stub ...
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Gorgonops
''Gorgonops'' (from el, Γοργών 'Gorgon' and 'eye, face', literally 'Gorgon eye' or 'Gorgon face') is an extinct genus of gorgonopsian therapsids, of which it is the type genus, having lived during the Late Permian (Wuchiapingian), about 260–254 million years ago in what is now South Africa. Despite its popularity, ''Gorgonops'' is a medium-sized gorgonopsian (about long maximum), regularly confused by the general public with the more massive ''Inostrancevia'', known from Russia, due to their similar appearance and the various media that tend to refer them by the name of the group they belong rather than by their genus names, which does not help in differentiation. History of discovery The holotype of the type species, ''Gorgonops torvus'', was in 1876 one of the first therapsids described, by Richard Owen, who also coined the name "Dinosauria" on the basis of the first known dinosaur fossils. It was also used as the type for which Richard Lydekker described the family ...
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Biarmosuchus
''Biarmosuchus'' is an extinct genus of biarmosuchian therapsids that lived around 267 mya during the Middle Permian period. ''Biarmosuchus'' was discovered in the Perm region of Russia. The first specimen was found in channel sandstone that was deposited by flood waters originating from the young Ural Mountains. Description ''Biarmosuchus'' was a medium-sized predator, similar in size to a large dog, grew up to 1.5–2 m in length with a skull length 15 cm (immature) to 21 cm. It was a lightly built, probably agile animal that would have fed on smaller tetrapods. Their legs are quite long, and the animals were probably quite agile in spite of their size. A large opening for the eye and a small temple opening common in primitive stem-mammals, this lends to a weak bite but how it ate is pure speculation. The teeth contained eight small incisors on the palate, followed by a canine tooth and a further five canine teeth. So together the species contained fourteen upper t ...
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Lycosuchus
''Lycosuchus'' ("wolf crocodile") is an extinct genus of carnivorous therocephalians which lived in the Middle Permian 265—260 Ma existing for approximately . As a member of the Lycosuchidae, the genus represents one of the earliest diverging therocephalians. The type species ''Lycosuchus vanderrieti'' is known from a handful of well-preserved specimens featuring the cranium and lower jaw; the holotype US D173 housed at the University of Stellenbosch, South Africa, is a near complete occluded skull. Specimen MB.R. 995, housed at the Museum für Naturkunde Berlin, Germany, consists of a near complete and isolated lower jaw, along with a partial snout and brain case. With the help of μCT data, Pusch et al (2020) described the endocranial anatomy of ''Lycosuchus vanderrieti''. It was a medium-sized predator, reaching 1.2 m (3.8 ft) in length with a skull 23 cm long., typical of early therocephalians. ''L. vanderrieti'' bore two functional canines in each maxilla, ...
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Raranimus
''Raranimus'' is an extinct genus of therapsids of the Middle Permian. It was described in 2009 from a partial skull found in 1998 from the Dashankou locality of the Xidagou Formation, outcropping in the Qilian Mountains of Gansu, China. The genus is the most basal known member of the clade Therapsida, to which the later Mammalia belong. Description ''Raranimus'' shares a number of features with later therapsids and ancestral Sphenacodontia. The skull consists of a well preserved rostrum. The teeth suggest a carnivorous lifestyle for ''Raranimus'', as the incisors are recurved and the second canines are serrated on their posterior edges. The incisors are morphologically similar to those seen in more derived theriodonts. The presence of two linguo−labially compressed canines is a diagnostic feature of ''Raranimus''. The presence of two functional canines is characteristic of sphenacodontids, and this condition is seen in no other therapsid other than ''Rananimus''. However, the ...
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Tetraceratops
''Tetraceratops insignis'' ("four-horned face emblem") is an extinct synapsid from the Early Permian that was formerly considered the earliest known representative of Therapsida, a group that includes mammals and their close extinct relatives. It is known from a single skull, discovered in Texas in 1908. According to a 2020 study, it should be classified as a primitive non-therapsid sphenacodont rather than a genuine basal therapsid. Description ''Tetraceratops'' is known from a single skull discovered in Texas in the early 1900s. Contrary to its genus name, ''Tetraceratops'' actually has six horns, one pair being on the premaxilla bones, one pair on the prefrontal bones, and one pair on the angular processes of the mandible. When it was discovered and described in 1908, the skull was still embedded in a matrix, and only the premaxilla and prefrontal pairs were visible. In life, thus, it would have resembled a large lizard with four horns on its snout, and a pair of large s ...
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