Allokotosaur
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Allokotosaur
Allokotosauria is a clade of early archosauromorph reptiles from the Middle Triassic, Middle to Late Triassic known from Asia, Africa, North America and Europe. Allokotosauria was first described and named when a new monophyletic grouping of specialized herbivorous archosauromorphs was recovered by Sterling J. Nesbitt, John J. Flynn, Adam C. Pritchard, J. Michael Parrish, Lovasoa Ranivoharimanana and André R. Wyss in 2015 in paleontology, 2015. The name Allokotosauria is derived from Greek language, Greek meaning "strange reptiles" in reference to unexpected grouping of early archosauromorph with a high disparity of features typically associated with herbivore, herbivory. History Nesbitt ''et al.'' (2015) defined the group as a stem-based taxon containing ''Azendohsaurus madagaskarensis'' and ''Trilophosaurus buettneri'' and all taxa more closely related to them than to ''Tanystropheus longobardicus'', ''Proterosuchus fergusi'', ''Protorosaurus speneri'' or ''Rhynchosaurus art ...
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Azendohsaurus Madagaskarensis
''Azendohsaurus'' is an extinct genus of herbivorous archosauromorph reptile from roughly the late Middle to early Late Triassic Period of Morocco and Madagascar. The type species, ''Azendohsaurus laaroussii'', was described and named by Jean-Michel Dutuit in 1972 based on partial jaw fragments and some teeth from Morocco. A second species from Madagascar, ''A. madagaskarensis'', was first described in 2010 by John J. Flynn and colleagues from a multitude of specimens representing almost the entire skeleton. The generic name "Azendoh lizard" is for the village of Azendoh, a local village near where it was first discovered in the Atlas Mountains. It was a bulky quadruped that unlike other early archosauromorphs had a relatively short tail and robust limbs that were held in an odd mix of sprawled hind limbs and raised forelimbs. It had a long neck and a proportionately small head with remarkably sauropod-like jaws and teeth. ''Azendohsaurus'' used to be classified as a herbivo ...
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Azendohsaurus
''Azendohsaurus'' is an extinct genus of herbivorous archosauromorph reptile from roughly the late Middle to early Late Triassic Period of Morocco and Madagascar. The type species, ''Azendohsaurus laaroussii'', was described and named by Jean-Michel Dutuit in 1972 based on partial jaw fragments and some teeth from Morocco. A second species from Madagascar, ''A. madagaskarensis'', was first described in 2010 by John J. Flynn and colleagues from a multitude of specimens representing almost the entire skeleton. The generic name "Azendoh lizard" is for the village of Azendoh, a local village near where it was first discovered in the Atlas Mountains. It was a bulky quadruped that unlike other early archosauromorphs had a relatively short tail and robust limbs that were held in an odd mix of sprawled hind limbs and raised forelimbs. It had a long neck and a proportionately small head with remarkably sauropod-like jaws and teeth. ''Azendohsaurus'' used to be classified as a herbivorous ...
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Azendohsauridae
Azendohsauridae is a family of allokotosaurian archosauromorphs that lived during the Middle to Late Triassic period, around 242-216 million years ago. The family was originally named solely for the eponymous ''Azendohsaurus'', marking out its distinctiveness from other allokotosaurs, but the family now includes four other genera: the basal genus '' Pamelaria'', the large horned herbivore '' Shringasaurus'', and two carnivorous genera grouped into the subfamily-level subclade Malerisaurinae, '' Malerisaurus'' and '' Puercosuchus'', and potentially also the dubious genus ''Otischalkia''. Most fossils of azendohsaurids have a Gondwanan distribution, with multiple species known across Morocco and Madagascar in Africa as well as India, although fossils of malerisaurine azendohsaurids have also been found in the southwestern United States of North America. Azendohsaurids are notable for the various dinosaur-like traits found in some species, including the sauropodomorph-like nec ...
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Archosauromorph
Archosauromorpha (Greek for "ruling lizard forms") is a clade of diapsid reptiles containing all reptiles more closely related to archosaurs (such as crocodilians and dinosaurs, including birds) rather than lepidosaurs (such as tuataras, lizards, and snakes). Archosauromorphs first appeared during the late Middle Permian or Late Permian, though they became much more common and diverse during the Triassic period. Although Archosauromorpha was first named in 1946, its membership did not become well-established until the 1980s. Currently Archosauromorpha encompasses four main groups of reptiles: the stocky, herbivorous allokotosaurs and rhynchosaurs, the hugely diverse Archosauriformes, and a polyphyletic grouping of various long-necked reptiles including ''Protorosaurus'', tanystropheids, and ''Prolacerta''. Other groups including pantestudines (turtles and their extinct relatives) and the semiaquatic choristoderes have also been placed in Archosauromorpha by some authors. A ...
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Pamelaria
''Pamelaria'' is an extinct genus of allokotosaurian archosauromorph reptile known from a single species, ''Pamelaria dolichotrachela'', from the Middle Triassic of India. ''Pamelaria'' has sprawling legs, a long neck, and a pointed skull with nostrils positioned at the very tip of the snout. Among early archosauromorphs, ''Pamelaria'' is most similar to ''Prolacerta'' from the Early Triassic of South Africa and Antarctica. Both have been placed in the family Prolacertidae. ''Pamelaria'', ''Prolacerta'', and various other Permo-Triassic reptiles such as ''Protorosaurus'' and ''Tanystropheus'' have often been placed in a group of archosauromorphs called Protorosauria (alternatively called Prolacertiformes), which was regarded as one of the most basal group of archosauromorphs. However, more recent phylogenetic analyses indicate that ''Pamelaria'' and ''Prolacerta'' are more closely related to Archosauriformes than are ''Protorosaurus'', ''Tanystropheus'', and other protorosaurs, ...
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Trilophosaurus Buettneri
''Trilophosaurus'' (Greek language, Greek for "lizard with three ridges") is a lizard-like trilophosaurid allokotosaur known from the Late Triassic of North America. It was a herbivore up to 2.5 m long. It had a short, unusually heavily built skull, equipped with massive, broad flattened cheek teeth with sharp shearing surfaces for cutting up tough plant material. Teeth are absent from the premaxilla and front of the lower jaw, which in life were probably equipped with a horny beak. The skull is also unusual in that the lower temporal fenestra, temporal opening is missing, giving the appearance of a euryapsida, euryapsid skull. Because of this, the trilophosaurs were once classified with placodonts within Sauropterygia. Robert L. Carroll, Carroll (1988) suggested that the lower opening may have been lost to strengthen the skull. ''Trilophosaurus'' is traditionally thought to include two valid species: the typical ''T. buettneri'' and the more robust ''T. jacobsi''. In 1993, p ...
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Trilophosauridae
Trilophosaurs are lizard-like Triassic allokotosaur reptiles related to the archosaurs. The best known genus is ''Trilophosaurus'', a herbivore up to long. It had a short, unusually heavily built skull, equipped with massive, broad flattened cheek teeth with sharp shearing surfaces for cutting up tough plant material. Teeth are absent from the premaxilla and front of the lower jaw, which in life were probably equipped with a horny beak. The skull is also unusual in that the lower temporal fenestra, temporal opening is missing, giving the appearance of a euryapsida, euryapsid skull, and originally the Trilophosaurs were classified with placodonts and sauropterygia. Carroll (1988) suggests that the lower opening may have been lost to strengthen the skull. Trilophosaurs are so far known only from the Late Triassic of North America and Europe. Below is a cladogram showing the phylogenetic relationships of Trilophosauridae within Archosauromorpha as recovered by Nesbitt ''et al.' ...
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Tanystropheus Longobardicus
''Tanystropheus'' (Greek ~ 'long' + 'hinged') is an extinct archosauromorph reptile from the Middle and Late Triassic epochs. It is recognisable by its extremely elongated neck, which measured long—longer than its body and tail combined. The neck was composed of 12–13 extremely elongated vertebrae. With its very long but relatively stiff neck, ''Tanystropheus'' has been often proposed and reconstructed as an aquatic or semi-aquatic reptile, a theory supported by the fact that the creature is most commonly found in semi-aquatic fossil sites wherein known terrestrial reptile remains are scarce. Fossils have been found in Europe. Complete skeletons of small individuals are common in the Besano Formation at Monte San Giorgio in Italy and Switzerland; other fossils have been found in the Middle East and China, dating from the Middle Triassic to the early part of the Late Triassic (Anisian, Ladinian, and Carnian stages).Dal Sasso, C. and Brillante, G. (2005). ''Dinosaurs of Italy ...
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Arctosaurus
''Arctosaurus'' is an extinct genus of archosauromorph, possibly an allokotosaurian, but was often classified as a sauropodomorph dinosaur between 1900 and 1976. Although it has also been classified as a theropod, recent review finds that the similarities it shares with theropods are spread throughout several groups of Late Triassic reptiles, and so it cannot be assigned any more specifically than to Archosauriformes. Other authors have suggested trilophosaurian affinities. Based on the size of the vertebra, a size of about in length is extrapolated. Discovery and naming It is based on holotype NMING: F14878, a neck vertebra that was found in 1859 by Captain Sherard Osborn on Cameron Island, Nunavut, Canada, in Late Triassic-age rocks of the Heiberg Formation The Heiberg Formation is a geological Formation (geology), formation in Northwest Territories, Canada.Weishampel, David B; et al. (2004). "Dinosaur distribution (Late Triassic, North America)." In: Weishampel, David B.; ...
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Protorosaurus Speneri
''Protorosaurus'' ("first lizard") is a genus of lizard-like early reptiles. Members of the genus lived during the late Permian period in what is now Germany and Great Britain. Once believed to have been an ancestor to lizards, ''Protorosaurus'' is now known to be one of the oldest and most primitive members of Archosauromorpha, the group that would eventually lead to archosaurs such as crocodilians and dinosaurs. Description ''Protorosaurus'' grew up to in length, and was a slender, lizard-like animal, vaguely resembling a monitor lizard, with long legs and a long neck. Discovery ''Protorosaurus'' was one of the first fossil reptiles to be described, being initially described in Latin in 1710 by from a specimen found in Thuringia in Germany, who considered the animal to be a crocodile, and most similar to the Nile crocodile (''C. niloticus''). Over a century later, in publications in 1830 and 1832 Hermann von Meyer recognised ''Protorosaurus'' as distinct extinct rep ...
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Reptile
Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians (tuatara). As of March 2022, the Reptile Database includes about 11,700 species. In the traditional Linnaean classification system, birds are considered a separate class to reptiles. However, crocodilians are more closely related to birds than they are to other living reptiles, and so modern cladistic classification systems include birds within Reptilia, redefining the term as a clade. Other cladistic definitions abandon the term reptile altogether in favor of the clade Sauropsida, which refers to all amniotes more closely related to modern reptiles than to mammals. The study of the traditional reptile orders, historically combined with that of modern amphibians, is called herpetology. The earliest known proto-reptiles originated around ...
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Proterosuchus Fergusi
''Proterosuchus'' is an extinct genus of archosauriform reptiles that lived during the Early Triassic. It contains three valid species: the type species ''P. fergusi'' and the referred species ''P. alexanderi'' and ''P. goweri''. All three species lived in what is now South Africa. The genus was named in 1903 by the South African paleontologist Robert Broom. The well-known genus ''Chasmatosaurus'' is a junior synonym of ''Proterosuchus''. ''Proterosuchus'' was a mid-sized quadrupedal reptile with a sprawling stance that could reach a length of up to . It had a large head and distinctively hooked snout. It was a predator, which may have hunted prey such as ''Lystrosaurus''. The lifestyle of ''Proterosuchus'' remains debated; it may have been terrestrial or it may have been a semiaquatic ambush predator similar to modern crocodiles. ''Proterosuchus'' is one of the earliest members of the clade Archosauriformes, which also includes crocodilians, pterosaurs, and dinosaurs, inc ...
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