Allochronic Speciation
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Allochronic Speciation
Allochronic speciation (also known as allochronic isolation, or temporal isolation) is a form of speciation (specifically ecological speciation) arising from reproductive isolation that occurs due to a change in breeding time that reduces or eliminates gene flow between two populations of a species. The term allochrony is used to describe the general ecological phenomenon of the differences in phenology that arise between two or more species—speciation caused by allochrony is effectively allochronic speciation. Environmental changes acting on a species population or populations can drive isolation. An important form of isolation is when populations are separated, not geographically, but temporally (by time). Genetic changes (mutations) over time can cause the two populations to differ—notably in phenology (events in a species life dictated by time such as breeding seasons); exhibiting unique phenotypes (the observable characteristics or traits of an organism). Scientists h ...
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Speciation
Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution within lineages. Charles Darwin was the first to describe the role of natural selection in speciation in his 1859 book ''On the Origin of Species''. He also identified sexual selection as a likely mechanism, but found it problematic. There are four geographic modes of speciation in nature, based on the extent to which speciating populations are isolated from one another: allopatric speciation, allopatric, peripatric speciation, peripatric, parapatric speciation, parapatric, and sympatric speciation, sympatric. Speciation may also be induced artificially, through animal husbandry, agriculture, or laboratory experiments of speciation, laboratory experiments. Whether genetic drift is a minor or major contributor to speciation is the subject of much ...
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Photoperiodism
Photoperiodism is the physiological reaction of organisms to the length of night or a dark period. It occurs in plants and animals. Plant photoperiodism can also be defined as the developmental responses of plants to the relative lengths of light and dark periods. They are classified under three groups according to the photoperiods: short-day plants, long-day plants, and day-neutral plants. Plants Many flowering plants (angiosperms) use a photoreceptor protein, such as phytochrome or cryptochrome, to sense seasonal changes in night length, or photoperiod, which they take as signals to flower. In a further subdivision, ''obligate'' photoperiodic plants absolutely require a long or short enough night before flowering, whereas ''facultative'' photoperiodic plants are more likely to flower under one condition. Phytochrome comes in two forms: Pr and Pfr. Red light (which is present during the day) converts phytochrome to its active form (pfr). This then triggers the plant to grow. ...
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Cline (biology)
In biology, a cline (from the Greek κλίνειν ''klinein'', meaning "to lean") is a measurable gradient in a single character (or biological trait) of a species across its geographical range. First coined by Julian Huxley in 1938, the "character" of the cline referred to is usually genetic (e.g. allele frequency, blood type), or phenotypic (e.g. body size, skin pigmentation). Clines can show smooth, continuous gradation in a character, or they may show more abrupt changes in the trait from one geographic region to the next. A cline refers to a spatial gradient in a specific, singular trait, rather than a collection of traits; a single population can therefore have as many clines as it has traits, at least in principle. Additionally, Huxley recognised that these multiple independent clines may not act in concordance with each other. For example, it has been observed that in Australia, birds generally become smaller the further towards the north of the country they are found ...
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Phenotype
In genetics, the phenotype () is the set of observable characteristics or traits of an organism. The term covers the organism's morphology or physical form and structure, its developmental processes, its biochemical and physiological properties, its behavior, and the products of behavior. An organism's phenotype results from two basic factors: the expression of an organism's genetic code, or its genotype, and the influence of environmental factors. Both factors may interact, further affecting phenotype. When two or more clearly different phenotypes exist in the same population of a species, the species is called polymorphic. A well-documented example of polymorphism is Labrador Retriever coloring; while the coat color depends on many genes, it is clearly seen in the environment as yellow, black, and brown. Richard Dawkins in 1978 and then again in his 1982 book ''The Extended Phenotype'' suggested that one can regard bird nests and other built structures such as cad ...
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Bamboo
Bamboos are a diverse group of evergreen perennial flowering plants making up the subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family. The origin of the word "bamboo" is uncertain, but it probably comes from the Dutch or Portuguese language, which originally borrowed it from Malay or Kannada. In bamboo, as in other grasses, the internodal regions of the stem are usually hollow and the vascular bundles in the cross-section are scattered throughout the stem instead of in a cylindrical arrangement. The dicotyledonous woody xylem is also absent. The absence of secondary growth wood causes the stems of monocots, including the palms and large bamboos, to be columnar rather than tapering. Bamboos include some of the fastest-growing plants in the world, due to a unique rhizome-dependent system. Certain species of bamboo can grow within a 24-hour period, at a rate of almost an hour (equivalent to 1 mm every 90 seco ...
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Periodical Cicadas
The term periodical cicada is commonly used to refer to any of the seven species of the genus ''Magicicada'' of eastern North America, the 13- and 17-year cicadas. They are called periodical because nearly all individuals in a local population are developmentally synchronized and emerge in the same year. Although they are sometimes called "locusts", this is a misnomer, as cicadas belong to the taxonomic order Hemiptera (true bugs), suborder Auchenorrhyncha, while locusts are grasshoppers belonging to the order Orthoptera. ''Magicicada'' belongs to the cicada tribe Lamotialnini, a group of genera with representatives in Australia, Africa, and Asia, as well as the Americas. ''Magicicada'' species spend around 99.5% of their long lives underground in an immature state called a nymph. While underground the nymphs feed on xylem fluids from the roots of deciduous forest trees in the eastern United States. In the spring of their 13th or 17th year mature cicada nymphs emerge between ...
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Sockeye Salmon
The sockeye salmon (''Oncorhynchus nerka''), also called red salmon, kokanee salmon, blueback salmon, or simply sockeye, is an anadromous species of salmon found in the Northern Pacific Ocean and rivers discharging into it. This species is a Pacific salmon that is primarily red in hue during spawning. They can grow up to in length and weigh . Juveniles remain in freshwater until they are ready to migrate to the ocean, over distances of up to . Their diet consists primarily of zooplankton. Sockeye salmon are semelparous, dying after they spawn. Some populations, referred to as kokanee, do not migrate to the ocean and live their entire lives in fresh water. Classification and name origin The sockeye salmon is the third-most common Pacific salmon species, after pink and chum salmon. ''Oncorhynchus'' comes from the Greek ὄγκος (onkos) meaning "barb", and ῥύγχος (rhynchos) meaning "snout". ''Nerka'' is the Russian name for the anadromous form. The name "sockeye" ...
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Salmon Run
''Salmon Run'' is a 1982 video game for the Atari 8-bit family created by Bill Williams and distributed via the Atari Program Exchange. ''Salmon Run'' was the first game in Williams's career, followed by a string of successes noted for their oddball concepts. The player takes the role of Sam the Salmon, swimming upriver to mate. Along the way he encounters waterfalls, a bear, fishermen, and seagulls. In 1983, ''Salmon Run'' was released for the VIC-20 by Synapse Software Synapse Software Corporation (marketed as SynSoft in the UK) was an American video game development and publishing company founded in 1981 by Ihor Wolosenko and Ken Grant. It initially focused on the Atari 8-bit family, then later developed for th ... under the name Showcase Software. Gameplay ''Salmon Run'' is an overhead view, vertically scrolling game. As the river scrolls, the player primarily movesside-to-side to avoid obstacles. Each player starts with one life and gains another for each successful ...
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Orbicella
''Orbicella'' is a genus of stony corals in the Merulinidae family. The Orbicella species complex comprises three sister species, namely ''Orbicella faveolata'', ''Orbicella annularis'' and ''Orbicella franksi'', all of which are shallow-water, zooxanthellate species and are native to the tropical western Atlantic Ocean, the Caribbean Sea and the Gulf of Mexico. These corals are ubiquitous and major reef-builders in the Caribbean. Their similar colony morphologies misled scientists to historically lump them into a single species, ''Montastraea annularis'', which included three morphotypes “bumpy”, “columnar” and “massive”. These growth forms were believed to arise as a response to abiotic factors (e.g., depth, light availability). This taxonomic classification was challenged by further ecological, reproductive, genetic, and morphologic evidence, which led to the re-description of three separate species, ''Montastraea faveolata'' (massive), ''M. annularis'' (columnar) ...
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Acropora
''Acropora'' is a genus of small polyp stony coral in the phylum Cnidaria. Some of its species are known as table coral, elkhorn coral, and staghorn coral. Over 149 species are described. ''Acropora'' species are some of the major reef corals responsible for building the immense calcium carbonate substructure that supports the thin living skin of a reef. Anatomy and distribution Depending on the species and location, ''Acropora'' species may grow as plates or slender or broad branches. Like other corals, ''Acropora'' corals are colonies of individual polyps, which are about 2 mm across and share tissue and a nerve net. The polyps can withdraw back into the coral in response to movement or disturbance by potential predators, but when undisturbed, they protrude slightly. The polyps typically extend further at night to help capture plankton and organic matter from the water. The species are distributed in the Indo-Pacific (over 100 species) and Caribbean (3 species). ...
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Reinforcement (speciation)
Reinforcement is a process of speciation where natural selection increases the reproductive isolation (further divided to pre-zygotic isolation and post-zygotic isolation) between two populations of species. This occurs as a result of selection acting against the production of hybrid individuals of low fitness. The idea was originally developed by Alfred Russel Wallace and is sometimes referred to as the Wallace effect. The modern concept of reinforcement originates from Theodosius Dobzhansky. He envisioned a species separated allopatrically, where during secondary contact the two populations mate, producing hybrids with lower fitness. Natural selection results from the hybrid's inability to produce viable offspring; thus members of one species who do not mate with members of the other have greater reproductive success. This favors the evolution of greater prezygotic isolation (differences in behavior or biology that inhibit formation of hybrid zygotes). Reinforcement is on ...
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