Agnostida
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Agnostida
Agnostida is an order of arthropod which have classically been seen as a group of highly modified trilobites, though some recent research has doubted this placement. Regardless, they appear to be close relatives as part of the Artiopoda. They are present in the Lower Cambrian fossil record along with trilobites from the Redlichiida, Corynexochida, and Ptychopariida orders, and were highly diverse throughout the Cambrian. Agnostidan diversity severely declined during the Cambrian-Ordovician transition, and the last agnostidans went extinct in the Late Ordovician. Systematics The Agnostida are divided into two suborders — Agnostina and Eodiscina — which are then subdivided into a number of families. As a group, agnostids are isopygous, meaning their pygidium is similar in size and shape to their cephalon. Most agnostid species were eyeless. The systematic position of the order Agnostida within the class Trilobita remains uncertain, and there has been continuing deb ...
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Ptychagnostidae
Ptychagnostidae is a family of agnostid trilobites from the 5th Stage to the Paibian Age of the Cambrian ( to million years ago). The family includes several important index fossils. Description Like all agnostids, ptychagnostids have cephalons and pygidia that are more or less uniform in size and shape ( isopygous). The thorax is composed of two body segments (somites). They are completely blind. Distribution and age range Ptychagnostidae has a cosmopolitan distribution. They existed during the 5th Stage of the Series 3 Epoch to the Paibian Age of the early Furongian Epoch in the Cambrian ( to million years ago). The earliest member of the family is '' Ptychagnostus praecurrens'' from the Burgess Shale fauna. Taxonomy Ptychagnostidae is classified under the superfamily Agnostoidea of the suborder Agnostina, order Agnostida. The family was first established by the Japanese paleontologist Teiichi Kobayashi in 1939. Its name comes from the type genus, ''Ptycha ...
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Peronopsidae
The Peronopsidae (which may also be called peronopsids) comprise the earliest family of the Agnostina suborder. Species of this family occurred on all paleocontinents. The earliest representatives of this family first occur just before the start of the Middle Cambrian, and the last disappeared just after the start of the Upper Cambrian. Distribution Peronopsidae are cosmopolitan. Temporal distribution Temporal distribution: * '' Archaeagnostus'', Toyonian ( ''Nephrolenellus multinodus''-zone) and Amgaian ('' Ovatoryctocara''-zone). * '' Eoagnostus'', Toyonian (''Nephrolenellus multinodus''-zone) and Amgaian (''Ovatoryctocara''-zone). * '' Peronopsis (Proacadagnostus)'', Amgaian (''Ovatoryctocara''- and lower ''Kounamkites''-zones). * '' Diplorrhina'', Amgaian (''Ovatoryctocara''- and ''Kounamkites''-zones) * '' Peronopsis (Peronopsis)'', Amgaian (''Ovatoryctocara''-, ''Kounamkites''- and ''Triplagnostus gibbus''-zones). * '' Pentagnostus (Pentagnostus)'', Amgaian (''Ovator ...
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Agnostidae
Agnostidae is a family of Agnostida trilobites. Like all Agnostina, they were eyeless and bore only two thoracic segments. They ranged in benthic waters across the globe from 508 to 461 million years ago, containing the following genera, among others: *'' Acmarhachis'' *''Agnostus ''Agnostus'' is a genus of agnostid trilobites, belonging to the family Agnostidae, that lived during the late Middle Cambrian – early Upper Cambrian (about 506 to 492 million years ago). It is the type genus of the family Agnostidae and is sub ...'' *'' Aistagnostus'' *'' Anglagnostus'' *'' Biciragnostus'' *'' Connagnostus'' *'' Distagnostus'' *'' Eolotagnostus'' *'' Gymnagnostus'' *'' Homagnostus'' *'' Idolagnostus'' *'' Innitagnostus'' *'' Ivshinagnostus'' *'' Kymagnostus'' *'' Lotagnostus'' *'' Micragnostus'' *'' Obelagnostus'' *'' Oncagnostus'' *'' Peronopsis'' *'' Phalacroma'' *'' Phalagnostus'' *'' Quadrahomagnostus'' *'' Raragnostus'' *'' Semagnostus'' *'' Strictagnostus'' *'' Trilobagnostu ...
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Ammagnostidae
Ammagnostidae is a family of trilobites in the suborder Agnostina, small, eyeless, isopygous trilobites with a thorax consisting of 2 segments only. Four genera have been assigned to it: ;''Ammagnostus'' Öpik, 1967 *'' Ammagnostus psammius'' Öpik, 1967 ( Type) *'' Ammagnostus bassus'' (Öpik, 1967) *'' Ammagnostus bella'' Guo & Luo, 1996 *'' Ammagnostus beltensis'' (Lochman, 1944) *'' Ammagnostus cryptus'' *'' Ammagnostus cylindratus'' Guo & Luo, 1996 *'' Ammagnostus duibianensis'' Lu & Lin, 1989 *'' Ammagnostus histus'' *'' Ammagnostus hunanensis'' *'' Ammagnostus integriceps'' Öpik, 1967 *'' Ammagnostus laiwuensis'' (Lorenz, 1906) *'' Ammagnostus mitis'' Öpik, 1967 *'' Ammagnostus sinensis'' Peng, 1987 *'' Ammagnostus wangcunensis'' Peng & Robison ;''Hadragnostus'' Öpik, 1967 *'' Hadragnostus las'' Öpik, 1967 ( Type) *'' Hadragnostus edax'' Fortey & Rushton, 1976 *'' Hadragnostus helixensis'' Jago & Cooper, 2005 *'' Hadragnostus modestus'' (Lochman, 1944) ;''Kormagnostu ...
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Yukoniidae
Yukoniidae S. Zhang, 1980 'nom. transl''. et ''emend''. Jell, ''in'' Whittington ''et al''., 1997 ''ex'' Yukoniinae S. Zhang in W. Zhang, Lu ''et al''., 1980is a small family of trilobites, belonging to the Eodiscina. Type Genus and species ''Yukonia intermedia'' Palmer 1968 (Plate 2, figures 14, 17–19, 22, 23, 27, 28; text figure 4). Taxonomy The Yukoniidae probably descended from the Tsunydiscidae and gave rise to the Eodiscidae. cited in Description Yukoniidae are typically isopygous, belonging to the Superfamily Eodiscoidea. The narrow glabella The glabella, in humans, is the area of skin between the eyebrows and above the nose. The term also refers to the underlying bone that is slightly depressed, and joins the two brow ridges. It is a cephalometric landmark that is just superior to ... is usually parallel sided, anteriorly rounded and separated from smooth anterior border by broad (''sag''.) preglabellar field which occupies about 25% of cephalic length e ...
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Hebediscidae
The Hebediscidae Kobayashi, 1944, are a family of trilobites belonging to the order Agnostida that lived during the Lower Cambrian (Atdabanian to Toyonian). They are small or very small, and have a thorax of two or three segments. The Hebediscidae include five genera (see box). Taxonomy The Hebediscidae probably descended from the Tsunydiscidae and gave rise to the Weymouthiidae. cited in Description Like all Agnostida, the Hebediscidae are diminutive and the headshield (or cephalon) and tailshield (or pygidium) are of approximately the same size (or isopygous) and outline. In the Hebediscidae, the central raised area of the cephalon (or glabella The glabella, in humans, is the area of skin between the eyebrows and above the nose. The term also refers to the underlying bone that is slightly depressed, and joins the two brow ridges. It is a cephalometric landmark that is just superior to ...) is wide at its rear end, has parallel sides or tapers forward, and without t ...
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Weymouthiidae
The Weymouthiidae Kobayashi 1943 are an extinct family of eodiscinid agnostid trilobites. They lived during the late Lower Cambrian and earliest Middle Cambrian (Botomian to Delamaran) in the so-called ''Olenellus''- and ''Eokochaspis''-zones in the former paleocontinents of Laurentia, Avalonia, Gondwana. The Weymouthiidae are all blind and lack free cheeks. Taxonomy Ancestors The Weymouthiidae have developed from a stock within the Hebediscidae The Hebediscidae Kobayashi, 1944, are a family of trilobites belonging to the order Agnostida that lived during the Lower Cambrian (Atdabanian to Toyonian). They are small or very small, and have a thorax of two or three segments. The Hebediscida .... Descendants The Weymouthiidae are a paraphyletic family because the Agnostina suborder is nested within it, particularly in the clade that further consists of the genera ''Mallagnostus'', ''Chelediscus'', ''Tannudiscus'' and ''Jinghediscus''. The trend in the Weymouthiid ...
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Trilobite
Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period () and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described. By the time trilobites first appeared in the fossil record, they were already highly diversified and geographically dispersed. Because trilobites had wide diversity and an easily fossilized exoskeleton, they left an extensive fossil record. The stu ...
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Calodiscidae
The Calodiscidae Kobayashi, 1943 'nom. transl.'' Öpik, 1975 ''ex'' Calodiscinae Kobayashi, 1943are a family of trilobites belonging to the order Agnostida that lived during the Lower Cambrian (Botomian and Toyonian). They are small or very small, and have a thorax of two or three segments. The Calodiscidae includes five genera (see box). Taxonomy The probable ancestors of the Calodiscidae are among the Tsunydiscidae. The Calodiscidae had no descendants. cited ''in'' Description Like all Agnostida, the Calodiscidae are diminutive and the headshield (or cephalon) and tailshield (or pygidium) are of approximately the same size (or isopygous) and outline. The central raised area of the cephalon (or glabella The glabella, in humans, is the area of skin between the eyebrows and above the nose. The term also refers to the underlying bone that is slightly depressed, and joins the two brow ridges. It is a cephalometric landmark that is just superior to ...) has parallel si ...
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Trilobites
Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period () and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described. By the time trilobites first appeared in the fossil record, they were already highly diversified and geographically dispersed. Because trilobites had wide diversity and an easily fossilized exoskeleton, they left an extensive fossil record. The stud ...
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Lower Cambrian
The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Cambrian biolo ...
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Artiopoda
The Artiopoda is a grouping of extinct arthropods that includes trilobites and their close relatives. It was erected by Hou and Bergström in 1997 to encompass a wide diversity of arthropods that would traditionally have been assigned to the Trilobitomorpha. Hou and Bergström used the name Lamellipedia as a superclass to replace Trilobitomorpha that was originally erected at the subphylum level, which they considered inappropriate. Trilobites, in part due to their mineralising exoskeletons, are by far the most diverse and long lived members of the clade, with most records of other members, which lack mineralised exoskeletons, being from Cambrian deposits. Description According to Stein and Selden (2012) artiopods are recognised by the possession of filiform antennulae, limbs with bilobate exopods, with the proximal lobe being elongate and bearing a lamella, while the distal lobe is paddle-shaped and setiforous (bearing hair-or bristle like structures). The limb endopod has se ...
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