Ptychagnostidae
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Ptychagnostidae
Ptychagnostidae is a family of agnostid trilobites from the 5th Stage to the Paibian Age of the Cambrian ( to million years ago). The family includes several important index fossils. Description Like all agnostids, ptychagnostids have cephalons and pygidia that are more or less uniform in size and shape ( isopygous). The thorax is composed of two body segments ( somites). They are completely blind. Distribution and age range Ptychagnostidae has a cosmopolitan distribution. They existed during the 5th Stage of the Series 3 Epoch to the Paibian Age of the early Furongian Epoch in the Cambrian ( to million years ago). The earliest member of the family is ''Ptychagnostus praecurrens'' from the Burgess Shale fauna. Taxonomy Ptychagnostidae is classified under the superfamily Agnostoidea of the suborder Agnostina, order Agnostida. The family was first established by the Japanese paleontologist Teiichi Kobayashi in 1939. Its name comes from the type genus, '' Pty ...
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Ptychagnostus Gibbus
''Ptychagnostus'' is a member of the agnostida that lived during the Cambrian period. Ptychagnostidae generally do not exceed one centimetre in length. Their remains are rarely found in empty tubes of the polychaete worm ''Selkirkia ''Selkirkia'' is a genus of predatory, tubicolous priapulid worms known from the Middle Cambrian Burgess Shale, Ogygopsis Shale and Puncoviscana Formation. 142 specimens of ''Selkirkia'' are known from the Greater Phyllopod bed, where they compr ...''. The genus probably ranged throughout the water column. It has two glabellar lobes, and three pygidial lobes. Type species ''Agnostus punctuosus'' Angelin, 1851 from the ''Pt. punctuosus'' Zone of the Alum Shale ( Drumian), Sweden (by original designation). Official ruling on the conservation of accepted usage of ''A. punctuosus'' as the type species was given by the International Commission on Zoological Nomenclature, 1993. Remarks ''Ptychagnostus affinis'' (Brøgger 1878) was once considered a sub ...
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Ptychagnostus Cuyanus
''Ptychagnostus'' is a member of the agnostida that lived during the Cambrian period. Ptychagnostidae generally do not exceed one centimetre in length. Their remains are rarely found in empty tubes of the polychaete worm ''Selkirkia ''Selkirkia'' is a genus of predatory, tubicolous priapulid worms known from the Middle Cambrian Burgess Shale, Ogygopsis Shale and Puncoviscana Formation. 142 specimens of ''Selkirkia'' are known from the Greater Phyllopod bed, where they compr ...''. The genus probably ranged throughout the water column. It has two glabellar lobes, and three pygidial lobes. Type species ''Agnostus punctuosus'' Angelin, 1851 from the ''Pt. punctuosus'' Zone of the Alum Shale ( Drumian), Sweden (by original designation). Official ruling on the conservation of accepted usage of ''A. punctuosus'' as the type species was given by the International Commission on Zoological Nomenclature, 1993. Remarks ''Ptychagnostus affinis'' (Brøgger 1878) was once considered a sub ...
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Ptychagnostus
''Ptychagnostus'' is a member of the agnostida that lived during the Cambrian period. Ptychagnostidae generally do not exceed one centimetre in length. Their remains are rarely found in empty tubes of the polychaete worm ''Selkirkia ''Selkirkia'' is a genus of predatory, tubicolous priapulid worms known from the Middle Cambrian Burgess Shale, Ogygopsis Shale and Puncoviscana Formation. 142 specimens of ''Selkirkia'' are known from the Greater Phyllopod bed, where they compr ...''. The genus probably ranged throughout the water column. It has two glabellar lobes, and three pygidial lobes. Type species ''Agnostus punctuosus'' Angelin, 1851 from the ''Pt. punctuosus'' Zone of the Alum Shale ( Drumian), Sweden (by original designation). Official ruling on the conservation of accepted usage of ''A. punctuosus'' as the type species was given by the International Commission on Zoological Nomenclature, 1993. Remarks ''Ptychagnostus affinis'' (Brøgger 1878) was once considered a sub ...
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Ptychagnostus Praecurrens
''Ptychagnostus'' is a member of the agnostida that lived during the Cambrian period. Ptychagnostidae generally do not exceed one centimetre in length. Their remains are rarely found in empty tubes of the polychaete worm ''Selkirkia''. The genus probably ranged throughout the water column. It has two glabellar lobes, and three pygidial lobes. Type species ''Agnostus punctuosus'' Angelin, 1851 from the ''Pt. punctuosus'' Zone of the Alum Shale (Drumian), Sweden (by original designation). Official ruling on the conservation of accepted usage of ''A. punctuosus'' as the type species was given by the International Commission on Zoological Nomenclature, 1993. Remarks ''Ptychagnostus affinis'' (Brøgger 1878) was once considered a subspecies of ''Pt. punctuosus''. Laurie (2008) grouped ''punctuosus'' and ''affinis'' within ''Ptychagnostus'', but preferred to place the closely related ''atavus'' within ''Acidusus''. Ptychagnostidae Genera Høyberget & Bruton (2008) HØYBERGET M. & ...
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Agnostida
Agnostida is an order of arthropod which have classically been seen as a group of highly modified trilobites, though some recent research has doubted this placement. Regardless, they appear to be close relatives as part of the Artiopoda. They are present in the Lower Cambrian fossil record along with trilobites from the Redlichiida, Corynexochida, and Ptychopariida orders, and were highly diverse throughout the Cambrian. Agnostidan diversity severely declined during the Cambrian-Ordovician transition, and the last agnostidans went extinct in the Late Ordovician. Systematics The Agnostida are divided into two suborders — Agnostina and Eodiscina — which are then subdivided into a number of families. As a group, agnostids are isopygous, meaning their pygidium is similar in size and shape to their cephalon. Most agnostid species were eyeless. The systematic position of the order Agnostida within the class Trilobita remains uncertain, and there has been continuing deb ...
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Wuliuan
The Wuliuan stage is the fifth stage of the Cambrian, and the first stage of the Miaolingian Series of the Cambrian. It was formally defined by the ICS in 2018. Its base is defined by the first appearance of the trilobite species ''Oryctocephalus indicus''; it ends with the beginning of the Drumian Stage, marked by the first appearance of the trilobite ''Ptychagnostus atavus'' around million years ago. The 'golden spike' that formally defines the base of the period is driven into the Wuliu-Zengjiayan(乌溜-曾家崖)section of the Kaili formation, near Balang Village in the Miaoling Mountains, Guizhou, China. GSSP Three sections were discussed as GSSP candidates: the Wuliu-Zengjiayan section near Balang in Guizhou province (China), a section on Split Mountain in Nevada (USA) and the "Molodo river section" along the Molodo river (Sakha Republic, Russia). The Wuliu-Zengjiayan section is an outcrop of the Kaili Formation in the Wuliu quarry. The first candidate for the beginn ...
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Superfamily (biology)
In biological classification, taxonomic rank is the relative level of a group of organisms (a taxon) in an ancestral or hereditary hierarchy. A common system consists of species, genus, family, order, class, phylum, kingdom, domain. While older approaches to taxonomic classification were phenomenological, forming groups on the basis of similarities in appearance, organic structure and behaviour, methods based on genetic analysis have opened the road to cladistics. A given rank subsumes under it less general categories, that is, more specific descriptions of life forms. Above it, each rank is classified within more general categories of organisms and groups of organisms related to each other through inheritance of traits or features from common ancestors. The rank of any ''species'' and the description of its ''genus'' is ''basic''; which means that to identify a particular organism, it is usually not necessary to specify ranks other than these first two. Consider a particular ...
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Epoch (geology)
The geologic time scale, or geological time scale, (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochronology (scientific branch of geology that aims to determine the age of rocks). It is used primarily by Earth scientists (including geologists, paleontologists, geophysicists, geochemists, and paleoclimatologists) to describe the timing and relationships of events in geologic history. The time scale has been developed through the study of rock layers and the observation of their relationships and identifying features such as lithologies, paleomagnetic properties, and fossils. The definition of standardized international units of geologic time is the responsibility of the International Commission on Stratigraphy (ICS), a constituent body of the International Union of Geological Sciences (IUGS), whose primary objective is to precisely define gl ...
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Furongian
The Furongian is the fourth and final epoch and series of the Cambrian. It lasted from to million years ago. It succeeds the Miaolingian series of the Cambrian and precedes the Lower Ordovician Tremadocian Stage. It is subdivided into three stages: the Paibian, Jiangshanian and the unnamed 10th stage of the Cambrian. Naming The Furongian was also known as the Cambrian Series 4, and the name replaced the older term Upper Cambrian and equivalent to the local term Hunanian. The present name was ratified by the International Commission on Stratigraphy in 2003. () means ' lotus' in Mandarin and refers to Hunan which is known as the "lotus state". Definition The lower boundary is defined in the same way as the GSSP of the Paibian Stage. Both begin with the first appearance of the trilobite ''Glyptagnostus reticulatus'' around million years ago. The upper boundary is the lower boundary and GSSP of the Tremadocian Stage which is the first appearance of the conodont ''Iapetognathus f ...
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Burgess Shale Fauna
A number of assemblages bear fossil assemblages similar in character to that of the Burgess Shale. While many are also preserved in a similar fashion to the Burgess Shale, the term "Burgess Shale-type fauna" covers assemblages based on taxonomic criteria only. Extent The fauna of the middle Cambrian has a cosmopolitan range. All assemblages preserving soft-part anatomy have a very similar fauna, even though they span almost every continent. The wide distribution has been attributed to the advent of pelagic larvae. Composition The fauna is composed of a range of soft-bodied organisms; creatures with hard, mineralised skeletons are rare, although trilobites are quite commonly found. The major soft-bodied groups are sponges, palaeoscolecid worms, lobopods, arthropods and anomalocaridids. Assemblages are typically diverse, with the most famous localities each containing in the region of 150 described species. The fauna of the Burgess Shale lived in the photic zone, as bottom-dwellin ...
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