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Aptery
Aptery is the anatomical condition of an animal completely lacking any kind of wings. An animal with this condition is said to be apterous. Most animal species belong to and are phylogenetic descendants of apterous taxa. These groups are said to be primarily apterous. Insects that are primarily apterous belong to the subclass Apterygota. Apterous is an adjective that means that the insect or organism is wingless and usually refers to a particular form of an insect that may have wings, or a wingless species in a group that typically has wings, e.g. many Orthoptera (grasshoppers and allies) and Hymenoptera (wasps). In some groups, one sex may be apterous while the other is winged, e.g. Mutillidae (velvet ants). In other cases a particular form of an insect (but not all individuals) will be apterous, e.g. some Tetrigidae (pygmy grasshoppers). Wingless animals which belong to or are phylogenetic descendants of winged taxa are said to be secondarily apterous. 5% of extant Pterygota are ...
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Velvet Ant
The Mutillidae are a family of more than 7,000 species of wasps whose wingless females resemble large, hairy ants. Their common name velvet ant refers to their dense pile of hair, which most often is bright scarlet or orange, but may also be black, white, silver, or gold. Their bright colors serve as aposematic signals. They are known for their extremely painful stings, (the sting of the species '' Dasymutilla klugii'' rated a 3 on the Schmidt pain index and lasts up to 30 minutes), and has resulted in the common name "cow killer" or "cow ant" being applied to the species '' Dasymutilla occidentalis.'' However, mutillids are not aggressive and sting only in defense. In addition, the actual toxicity of their venom is much lower than that of honey bees or harvester ants. Unlike true ants, they are solitary, and lack complex social systems. Distribution Mutillidae can be found worldwide with about 230 genera or subgenera and around 8,000 species worldwide. Over 400 species occ ...
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Wing
A wing is a type of fin that produces lift while moving through air or some other fluid. Accordingly, wings have streamlined cross-sections that are subject to aerodynamic forces and act as airfoils. A wing's aerodynamic efficiency is expressed as its lift-to-drag ratio. The lift a wing generates at a given speed and angle of attack can be one to two orders of magnitude greater than the total drag on the wing. A high lift-to-drag ratio requires a significantly smaller thrust to propel the wings through the air at sufficient lift. Lifting structures used in water include various foils, such as hydrofoils. Hydrodynamics is the governing science, rather than aerodynamics. Applications of underwater foils occur in hydroplanes, sailboats and submarines. Etymology and usage For many centuries, the word "wing", from the Old Norse ''vængr'', referred mainly to the foremost limbs of birds (in addition to the architectural aisle). But in recent centuries the word's meaning has ...
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Phylogenetic
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does ...
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Taxon
In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and given a particular ranking, especially if and when it is accepted or becomes established. It is very common, however, for taxonomists to remain at odds over what belongs to a taxon and the criteria used for inclusion. If a taxon is given a formal scientific name, its use is then governed by one of the nomenclature codes specifying which scientific name is correct for a particular grouping. Initial attempts at classifying and ordering organisms (plants and animals) were set forth in Carl Linnaeus's Linnaean taxonomy, system in ''Systema Naturae'', 10th edition (1758), as well as an unpublished work by Bernard de Jussieu, Bernard and Antoine Laurent de Jussieu. The idea of a unit-based system of biological classification was first mad ...
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Apterygota
The name Apterygota is sometimes applied to a subclass of small, agile insects, distinguished from other insects by their lack of wings in the present and in their evolutionary history; notable examples are the silverfish, the firebrat, and the jumping bristletails. Their first known occurrence in the fossil record is during the Devonian period, 417–354 million years ago. The group Apterygota is not a clade; it is paraphyletic, and not universally recognized. As defined, the group contains two separate clades of wingless insects: Archaeognatha comprises jumping bristletails, while Zygentoma comprises silverfish and firebrats. The Zygentoma are in the clade Dicondylia with winged insects, a clade that includes all other insects, while Archaeognatha is sister to this lineage.A. Blanke, M. Koch, B. Wipfler, F. Wilde, B. Misof (2014) Head morphology of ''Tricholepidion gertschi'' indicates monophyletic Zygentoma. Frontiers in Zoology 11:16 doi:10.1186/1742-9994-11-16 The nymphs ...
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Orthoptera
Orthoptera () is an order of insects that comprises the grasshoppers, locusts, and crickets, including closely related insects, such as the bush crickets or katydids and wētā. The order is subdivided into two suborders: Caelifera – grasshoppers, locusts, and close relatives; and Ensifera – crickets and close relatives. More than 20,000 species are distributed worldwide. The insects in the order have incomplete metamorphosis, and produce sound (known as a "stridulation") by rubbing their wings against each other or their legs, the wings or legs containing rows of corrugated bumps. The tympanum, or ear, is located in the front tibia in crickets, mole crickets, and bush crickets or katydids, and on the first abdominal segment in the grasshoppers and locusts. These organisms use vibrations to locate other individuals. Grasshoppers and other orthopterans are able to fold their wings (i.e. they are members of Neoptera). Etymology The name is derived from the Greek ὀρθό ...
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Hymenoptera
Hymenoptera is a large order (biology), order of insects, comprising the sawfly, sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are Parasitoid wasp, parasitic. Females typically have a special ovipositor for inserting eggs into hosts or places that are otherwise inaccessible. This ovipositor is often modified into a stinger. The young develop through holometabolism (complete metamorphosis (biology), metamorphosis)—that is, they have a wormlike larval stage and an inactive pupal stage before they mature. Etymology The name Hymenoptera refers to the wings of the insects, but the original derivation is ambiguous. All references agree that the derivation involves the Ancient Greek language, Ancient Greek wikt:πτερόν, πτερόν (''pteron'') for wing. The Ancient Greek wikt:ὑμήν, ὑμήν (''hymen'') for membrane provides a plausible etymology for the term bec ...
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Mutillidae
The Mutillidae are a family of more than 7,000 species of wasps whose wingless females resemble large, hairy ants. Their common name velvet ant refers to their dense pile of hair, which most often is bright scarlet or orange, but may also be black, white, silver, or gold. Their bright colors serve as aposematic signals. They are known for their extremely painful stings, (the sting of the species ''Dasymutilla klugii'' rated a 3 on the Schmidt pain index and lasts up to 30 minutes), and has resulted in the common name "cow killer" or "cow ant" being applied to the species '' Dasymutilla occidentalis.'' However, mutillids are not aggressive and sting only in defense. In addition, the actual toxicity of their venom is much lower than that of honey bees or harvester ants. Unlike true ants, they are solitary, and lack complex social systems. Distribution Mutillidae can be found worldwide with about 230 genera or subgenera and around 8,000 species worldwide. Over 400 species occur ...
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Tetrigidae
Tetrigidae is an ancient family in the order Orthoptera, which also includes similar families such as crickets, grasshoppers, and their allies. Species within the Tetrigidae are variously called groundhoppers, pygmy grasshoppers,Borror DJ, Tripplehorn CA, Johnson NF (1989) An Introduction to the Study of Insects, 6th edition. Harcourt Brace College Publishers. New York. pg 213 pygmy devils or (mostly historical) "grouse locusts".Imms AD, rev. Richards OW & Davies RG (1970) ''A General Textbook of Entomology'' 9th Ed. Methuen 886 pp. Diagnostic characteristics Tetrigidae are typically less than 20 mm in length and are recognizable by a long pronotum. This pronotum extends over the length of the abdomen, sometimes to the tip of the wings, and ends in a point. In other Orthoptera, the pronotum is short and covers neither the abdomen nor the wings. Tetrigidae are generally cryptic in coloration. Some species have enlarged pronota that mimic leaves, stones or twigs. Other chara ...
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Pterygota
The Pterygota ( grc, πτερυγωτός, pterugōtós, winged) are a subclass of insects that includes the winged insects. It also includes insect orders that are secondarily wingless (that is, insect groups whose ancestors once had wings but that have lost them as a result of subsequent evolution). The pterygotan group comprises almost all insects. The insect orders not included are the Archaeognatha (jumping bristletails) and the Zygentoma (silverfishes and firebrats), two primitively wingless insect orders. Also not included is Entognatha, which consist of three orders no longer considered to be insects: Protura, Collembola, and Diplura. Systematics Traditionally, this group was divided into the infraclasses Paleoptera and Neoptera. The former are nowadays strongly suspected of being paraphyletic, and better treatments (such as dividing or dissolving the group) are presently being discussed. In addition, it is not clear how exactly the neopterans are related among each o ...
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Fleas
Flea, the common name for the order Siphonaptera, includes 2,500 species of small flightless insects that live as external parasites of mammals and birds. Fleas live by ingesting the blood of their hosts. Adult fleas grow to about long, are usually brown, and have bodies that are "flattened" sideways or narrow, enabling them to move through their hosts' fur or feathers. They lack wings; their hind legs are extremely well adapted for jumping. Their claws keep them from being dislodged, and their mouthparts are adapted for piercing skin and sucking blood. They can leap 50 times their body length, a feat second only to jumps made by another group of insects, the superfamily of froghoppers. Flea larvae are worm-like, with no limbs; they have chewing mouthparts and feed on organic debris left on their hosts' skin. Genetic evidence indicates that fleas are a specialised lineage of parasitic scorpionflies (Mecoptera) ''sensu lato'', most closely related to the family Nannochorist ...
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Notoptera
The wingless insect order Notoptera, a group first proposed in 1915, had been largely unrecognized since its original conception, until resurrected in 2004. As now defined, the order comprises five families, three of them known only from fossils, two known from both fossil and living representatives, and fewer than 60 known species in total. History of research The name was originally coined in 1915 for a group of fossil orthopteroids, and largely forgotten until it was resurrected and redefined ("Notoptera Crampton ''sensu novum''") by Engel and Grimaldi in 2004 (after the discovery of living Mantophasmatidae), who recommended to give a single order that includes both the living and fossil representatives of the lineage. Terry and Whiting in 2005 independently proposed a new name, "Xenonomia", for the same lineage of insects (including the Grylloblattodea and Mantophasmatodea). In 2006, Cameron, Barker, and Whiting studied the mitochondrial genomics of the Mantophasmatodea, ...
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