Ancient Whales
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Ancient Whales
Archaeoceti ("ancient whales"), or Zeuglodontes in older literature, is a paraphyletic group of primitive cetaceans that lived from the Early Eocene to the late Oligocene (). Representing the earliest cetacean radiation, they include the initial amphibious stages in cetacean evolution, thus are the ancestors of both modern cetacean suborders, Mysticeti and Odontoceti. This initial diversification occurred in the shallow waters that separated India and Asia , resulting in some 30 species adapted to a fully oceanic life. Echolocation and filter-feeding evolved during a second radiation . All archaeocetes from the Ypresian (56–47.8 mya) and most from the Lutetian (47.8–41.3 mya) are known exclusively from Indo-Pakistan, but Bartonian (41.3–38.0 mya) and Priabonian (38.0–33.9 mya) genera are known from across Earth, including North America, Egypt, New Zealand, and Europe. Although no consensus exists regarding the mode of locomotion of which cetaceans were capable during t ...
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William Henry Flower
Sir William Henry Flower (30 November 18311 July 1899) was an English surgeon, museum curator and comparative anatomist, who became a leading authority on mammals and especially on the primate brain. He supported Thomas Henry Huxley in an important controversy with Richard Owen about the human brain and eventually succeeded Owen as Director of the Natural History Museum in London. Origins Born on 30 November 1831 in his father's house at Stratford-upon-Avon in Warwickshire, he was the second son of Selina née Greaves (d. 1884), eldest daughter of Mary Whitehead and John Greaves, and Edward Fordham Flower, founder of the town brewery. His grandfather Richard Flower had married Elizabeth Fordham and settled at Albion, Illinois, where his father grew up. His uncles included the slate entrepreneur John Whitehead Greaves and William Pickering, Governor of Washington. His elder brother Charles Edward Flower ran the family brewery with the third brother Edgar Flower, while he ch ...
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Kutchicetus Minimus
''Kutchicetus'' is an extinct genus of early whale of the family Remingtonocetidae that lived during Early-Middle Eocene (Lutetian and Ypresian) in what is now the coastal border of Pakistan and India (, paleocoordinates ). It is closely related to ''Andrewsiphius'' with which it was synonymized by . proposed a new clade, Andrewsiphiinae, for the two species. Later authors, however, still accept both as separate genera. ''Kutchicetus'' is smaller than other remingtonocetids, and probably is the smallest Eocene cetacean. With its extremely narrow snout, it resembles ''Remingtonocetus'' and ''Dalanistes'', but its strong tail distinguishes it from both ''Remingtonocetus'' and ''Andrewsiphius''. Its limbs were short. ''Kutchicetus''' vertebral formula is 7, 15, 8, 4, 20–25. Its four fused sacral vertebrae were probably articulated to the hip bone and the numerous tail vertebrae were robust and elongated in contrast to its short and relatively gracile limb bones. This mo ...
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Protocetidae
Protocetidae, the protocetids, form a diverse and heterogeneous group of extinct cetaceans known from Asia, Europe, Africa, South America, and North America. Description There were many genera, and some of these are very well known (e.g., ''Rodhocetus''). Known protocetids had large fore- and hindlimbs that could support the body on land, and it is likely that they lived amphibiously: in the sea and on land. It is unclear at present whether protocetids had flukes (the horizontal tail fin of modern cetaceans). However, what is clear is that they are adapted even further to an aquatic life-style. In ''Rodhocetus'', for example, the sacrum – a bone that in land-mammals is a fusion of five vertebrae that connects the pelvis with the rest of the vertebral column – was divided into loose vertebrae. However, the pelvis retain a sacroiliac joint. Furthermore, the nasal openings are now halfway up the snout; a first step towards the telescoped condition in modern whales. ...
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Synapomorphy
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis T ...
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Dorudon Atrox-lateral
''Dorudon'' ("spear-tooth") is a genus of extinct basilosaurid ancient whales that lived alongside ''Basilosaurus'' 40.4 to 33.9 million years ago in the Eocene. It was a small whale, with ''D. atrox'' measuring long and weighing . ''Dorudon'' lived in warm seas around the world and fed on small fish and mollusks. Fossils have been found along the former shorelines of the Tethys Sea in present-day Egypt and Pakistan, as well as in the United States, New Zealand, and Western Sahara. Taxonomic history described ''Dorudon serratus'' based on a fragmentary maxilla and a few teeth found in South Carolina. He concluded that the teeth must have belonged to a mammal since they were two-rooted, that they must have been teeth from a juvenile since they were hollow, and also noted their similarity to the teeth then described for ''Zeuglodon'' (''Basilosaurus''). When exploring the type locality, Gibbes discovered a lower jaw and twelve caudal vertebrae, which he felt obliged to assign ...
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Maiacetus
''Maiacetus'' ("mother whale") is a genus of early middle Eocene (c. 47.5 mya) cetacean from the Habib Rahi Formation of Pakistan. Paleobiology The genus contains a single species, ''Maiacetus inuus'', first described in 2009 on the basis of two specimens, including a specimen which has been interpreted as a pregnant female and its fetus. This represents the first description of a fetal skeleton of an archaeocete. The position of the fetus (head-first) suggests that the animal gave birth on land."Earliest whales gave birth on land"
'''', 3 February 2009
Modern whales generally give birth tail first, while land mammals give birth head first. ...
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Remingtonocetidae
Remingtonocetidae is a diverse family of early aquatic mammals of the order Cetacea. The family is named after paleocetologist Remington Kellogg. Description Remingtonocetids have long and narrow skulls with the external nare openings located on the front of the skull. Their frontal shields are narrow and their orbits small. Their mouth has a convex palate and an incompletely fused mandibular symphysis. The dental formula is . The anterior teeth are flattened mediolaterally, making them appear shark-like. In the postcranial skeleton, the cervical vertebrae are relatively long and the sacrum is composed of four vertebrae of which at least three are fused. The acetabular notch is narrow or closed and on the femoral head the fovea is absent. Cranial fossils are common but dental remains are rare. The postcrania morphology is based entirely on a single specimen of ''Kutchicetus'' which was small and had a long and muscular back and tail. Perhaps remintonocetids swam like t ...
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Himalayacetus
''Himalayacetus'' is an extinct genus of carnivorous aquatic mammal of the family Ambulocetidae. The holotype was found in Himachal Pradesh, India, (: paleocoordinates ) in what was the remnants of the ancient Tethys Ocean during the Early Eocene. This makes ''Himalayacetus'' the oldest archaeocete known, extending the fossil record of whales some 3.5 million years. ''Himalayacetus'' lived in the ancient coastline of the ancient Tethys Ocean before the Indian Plate had collided with the Cimmerian coast. Like ''Gandakasia'', ''Himalayacetus'' is only known from a single jaw fragment, making comparisons to other ambulocetids difficult. Description Upon its discovery, ''Himalayacetus'' was described as a pakicetid because the dentary has a small mandibular canal and a dentition similar to ''Pakicetus''. assigned ''Himalaycetus'' to the ambulocetids. Etymology ''Himalayacetus'' was named by . Its type is ''Himalayacetus subathuensis'' after the Himalayas, ''cetus'', "whal ...
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Osteosclerosis
Osteosclerosis is a disorder that is characterized by abnormal hardening of bone and an elevation in bone density. It may predominantly affect the medullary portion and/or cortex of bone. Plain radiographs are a valuable tool for detecting and classifying osteosclerotic disorders. It can manifest in localized or generalized osteosclerosis. Localized osteosclerosis can be caused by Legg–Calvé–Perthes disease, sickle-cell disease and osteoarthritis among others. Osteosclerosis can be classified in accordance with the causative factor into acquired and hereditary. Types Acquired osteosclerosis * Osteogenic bone metastasis caused by carcinoma of prostate and breast * Paget's disease of bone * Myelofibrosis (primary disorder or secondary to intoxication or malignancy) * Osteosclerosing types of chronic osteomyelitis * Hypervitaminosis D * hyperparathyroidism * Schnitzler syndrome * Mastocytosis Skeletal fluorosis * Monoclonal IgM Kappa cryoglobulinemia * Hepatitis C. Heredita ...
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Ambulocetidae
Ambulocetidae is a family of early cetaceans from Pakistan. The genus ''Ambulocetus'', after which the family is named, is by far the most complete and well-known ambulocetid genus due to the excavation of an 80% complete specimen of ''Ambulocetus natans''. The other two genera in the family, ''Gandakasia'' and ''Himalayacetus'', are known only from teeth and mandibular fragments. Retaining large hindlimbs, it was once thought that they could walk on land—indeed, their name means "walking whales"—, but recent research suggests they may have been fully aquatic like modern cetaceans. Description The most basal of marine cetaceans, ambulocetids lived in shallow near-shore environments such as estuaries and bays, but were still dependent on fresh water during some stage of their life. Some of the characteristics related to sound transmission found in the lower jaws of modern whales that were absent in pakicetids are present in ambulocetids. They probably swam by paddling their l ...
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Ossicles
The ossicles (also called auditory ossicles) are three bones in either middle ear that are among the smallest bones in the human body. They serve to transmit sounds from the air to the fluid-filled labyrinth (cochlea). The absence of the auditory ossicles would constitute a moderate-to-severe hearing loss. The term "ossicle" literally means "tiny bone". Though the term may refer to any small bone throughout the body, it typically refers to the malleus, incus, and stapes (hammer, anvil, and stirrup) of the middle ear. Structure The ossicles are, in order from the eardrum to the inner ear (from superficial to deep): the malleus, incus, and stapes, terms that in Latin are translated as "the hammer, anvil, and stirrup". * The malleus ( la, "hammer") articulates with the incus through the incudomalleolar joint and is attached to the tympanic membrane (eardrum), from which vibrational sound pressure motion is passed. * The incus ( la, "anvil") is connected to both the other bones. * ...
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Pachyostotic
Pachyostosis is a non-pathological condition in vertebrate animals in which the bones experience a thickening, generally caused by extra layers of lamellar bone. It often occurs together with bone densification (osteosclerosis), reducing inner cavities. This joint occurrence is called pachyosteosclerosis. However, especially in the older literature, "pachyostosis" is often used loosely, referring to all osseous specializations characterized by an increase in bone compactness and/or volume. It occurs in both terrestrial and, especially, aquatic or semi-aquatic vertebrates. In aquatic animals, such as seacows (manatees and dugongs), ''Thalassocnus'', and plesiosaurs, pachyostosis in the thoracic region provides (or provided) ballast against the air-filled lungs. This maintains neutral buoyancy in aquatic habitats. Most giant deer showed pronounced pachyostosis of the mandible and skull. It has been suggested that this served to store minerals for antler growth. Many Pachycephalosau ...
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