Zhenyuanopterus
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Zhenyuanopterus
''Zhenyuanopterus'' is a genus of boreopterid pterosaur which is known from Lower Cretaceous (early Aptian stage) Yixian Formation of Liaoning, China. It contains one species, ''Zhenyuanopterus longirostris'', which was first described and named by Lü Junchang. Discovery and naming ''Zhenyuanopterus longirostris'' was named by paleontologist Lü Junchang in 2005. The genus is named after Sun Zhenyuan, who gave Lü Junchang the fossils along with Zi Fan. The specific name ("long-snouted") refers to the long snout of the pterosaur. One specimen is known; a complete skeleton, it was found in the Huangbanjigou locality of the Yixian Formation, and it is catalogued as GLGMV 0001 in the Guilin Longshan Geological Museum in Guangxi, China. A second, larger partial specimen, consisting of the rear half of the body, was found at the same locality and catalogued as XHPM1088 at the Dalian Xinghai Paleontological Museum. In 2014, Teng Fangfang and colleagues described it. Description T ...
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Boreopterid
Boreopteridae (meaning "northern wings") is a group of pterodactyloid pterosaurs from the Aptian-age Lower Cretaceous Yixian Formation of Liaoning, China. Classification In 2006, Lü Junchang and colleagues named the clade Boreopteridae for the clade containing the common ancestor of ''Boreopterus'' and ''Feilongus'' and all its descendants, which the authors reclassified as close relatives of the ornithocheirids. (''Feilongus'' had originally been considered a gallodactylid). Many possible boreopterids were subsequently described, one possible example being ''Aetodactylus'', which has been claimed to be similar to ''Boreopterus''. Originally considered close relatives of the ornithocheirids, many of these supposed boreopterids have been found to belong to other groups of the pterodactyloid lineage. In 2012, a phylogenetic analysis by Lü ''et al.'' divided the Boreopteridae into two subfamilies: Boreopterinae, comprising ''Boreopterus'' and ''Zhenyuanopterus'', and Moganopterina ...
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Boreopterinae
Boreopteridae (meaning "northern wings") is a group of pterodactyloid pterosaurs from the Aptian-age Lower Cretaceous Yixian Formation of Liaoning, China. Classification In 2006, Lü Junchang and colleagues named the clade Boreopteridae for the clade containing the common ancestor of ''Boreopterus'' and ''Feilongus'' and all its descendants, which the authors reclassified as close relatives of the ornithocheirids. (''Feilongus'' had originally been considered a gallodactylid). Many possible boreopterids were subsequently described, one possible example being ''Aetodactylus'', which has been claimed to be similar to ''Boreopterus''. Originally considered close relatives of the ornithocheirids, many of these supposed boreopterids have been found to belong to other groups of the pterodactyloid lineage. In 2012, a phylogenetic analysis by Lü ''et al.'' divided the Boreopteridae into two subfamilies: Boreopterinae, comprising ''Boreopterus'' and ''Zhenyuanopterus'', and Moganopter ...
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Boreopteridae
Boreopteridae (meaning "northern wings") is a group of pterodactyloid pterosaurs from the Aptian-age Lower Cretaceous Yixian Formation of Liaoning, China. Classification In 2006, Lü Junchang and colleagues named the clade Boreopteridae for the clade containing the common ancestor of ''Boreopterus'' and ''Feilongus'' and all its descendants, which the authors reclassified as close relatives of the ornithocheirids. (''Feilongus'' had originally been considered a gallodactylid). Many possible boreopterids were subsequently described, one possible example being ''Aetodactylus'', which has been claimed to be similar to ''Boreopterus''. Originally considered close relatives of the ornithocheirids, many of these supposed boreopterids have been found to belong to other groups of the pterodactyloid lineage. In 2012, a phylogenetic analysis by Lü ''et al.'' divided the Boreopteridae into two subfamilies: Boreopterinae, comprising ''Boreopterus'' and ''Zhenyuanopterus'', and Moganopterina ...
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Boreopterus
''Boreopterus'' is a genus of boreopterid pterodactyloid pterosaur from the Barremian-Aptian-age Lower Cretaceous Yixian Formation of Dalian, Liaoning, China. Etymology ''Boreopterus'' was named in 2005 by Lü Junchang and Ji Qiang. The type species is ''Boreopterus cuiae''. The genus name is derived from Greek ''boreios'', "northern" and ''pteron'', "wing". The specific epithet honors Cui Xu. Description ''Boreopterus'' is based on holotype JZMP-04-07-3, a nearly complete but crushed skeleton and skull. The skull is 235 millimeters long (9.25 inches), low and elongated with a rounded tip. Its wingspan is estimated to have been around 1.45 meters (4.76 feet). Its teeth, especially the anterior nine pairs, are quite large, forming a mesh of sharp teeth at the front of the mouth; the third and fourth teeth from the front are the largest. There are at least 27 teeth in each side of both the upper and lower jaws, which is a large amount. Classification Lü and Ji ...
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Ludodactylus
''Ludodactylus'' (meaning "play finger") is a genus of anhanguerid pterodactyloid pterosaur from the Early Cretaceous period (Aptian stage) of what is now the Crato Formation of the Araripe Basin in Ceará, Brazil. The type and only species is ''L. sibbicki''. The generic name ''Ludodactylus'' refers to the fact that the animal had the combination of teeth and a ''Pteranodon''-like head crest, similar to many toy pterosaurs, and no such creature was known to exist until the discovery of ''Ludodactylus''. However, ''Ludodactylus'' is not the only pterosaur known to possess these features, its very close relative ''Caulkicephalus'' is another example. Etymology The genus was named by Eberhard Frey ''et al.'' in 2003 and contains one known species, ''Ludodactylus sibbicki''. The name is derived from Latin ''ludus'', "play" and Greek ''daktylos'', "finger". ''Ludus'' refers to the fact, long lamented by paleontologists, that many toy pterosaurs combined teeth with a ''Pteranodon''-l ...
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Anhangueria
Anhangueria (or anhanguerians) is a group of pterosaurs belonging to the clade Pteranodontoidea. Fossil remains of this group date back from the Early to Late Cretaceous periods (Valanginian to Turonian stages), around 140 to 92.5 million years ago. Anhangueria was named by paleontologists Taissa Rodrigues and Alexander Kellner in a review of ''Ornithocheirus'' species in 2013, they defined the clade as a branch-based taxon consisting of all pteranodontoids more closely related to ''Anhanguera blittersdorffi'' than to ''Istiodactylus latidens'' and ''Cimoliopterus cuvieri''. Classification Anhangueria originally only contained the genera '' Brasileodactylus'', ''Camposipterus'', ''Cearadactylus'', ''Ludodactylus'' as well as the family Anhangueridae, however, recent analyses had recovered the family Hamipteridae within this clade as well. In 2014, paleontologist Brian Andres and colleagues assigned more groups and genera within this clade, this included ''Guidraco'', the subfa ...
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Ornithocheiromorpha
Ornithocheiromorpha (from Ancient Greek, meaning "bird hand form") is a group of pterosaurs within the suborder Pterodactyloidea. Fossil remains of this group date back from the Early Cretaceous, Early to Late Cretaceous Period (geology), periods (Valanginian to Turonian stages), around 140 to 92.5 million years ago. Ornithocheiromorphs were discovered worldwide except Antarctica, though most genera were recovered in Europe, Asia and South America. They were the most diverse and successful pterosaurs during the Early Cretaceous, but throughout the Late Cretaceous they were replaced by better adapted and more advanced pterosaur species such the pteranodontids and azhdarchoids. The Ornithocheiromorpha was defined in 2014 by Andres and colleagues, and they made Ornithocheiromorpha the most inclusive clade containing ''Ornithocheirus'', but not ''Pteranodon''. Ornithocheiromorphs are considered to be some of the largest animals to have ever flown. Members of this group are also rega ...
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Lonchodectidae
Lonchodectidae or LonchodraconidaePêgas, R.V., Holgado, B., Leal, M.E.C., 2019. "''Targaryendraco wiedenrothi'' gen. nov. (Pterodactyloidea, Pteranodontoidea, Lanceodontia) and recognition of a new cosmopolitan lineage of Cretaceous toothed pterodactyloids", ''Historical Biology'', 1–15. doi:10.1080/08912963.2019.1690482 is a group of pterosaurs within the clade Pterodactyloidea. It has variously been considered to be within Ctenochasmatoidea, Azhdarchoidea and Pteranodontia.Witton, M.P., Martill, D.M., and Green, M. (2009). "On pterodactyloid diversity in the British Wealden (Lower Cretaceous) and a reappraisal of “Palaeornis” ''cliftii'' Mantell, 1844." ''Cretaceous Research'', 30: 676–686. They are notable for their high, conical tooth sockets and raised alveolar margins. Taxonomic history Lonchodectidae was first named by paleontologist Reginald Walter Hooley in 1914, and was first considered to only contain species of ''Lonchodectes''.Unwin, David M. 2001. "An over ...
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Guidraco
''Guidraco'' (Chin. ''gui'' (鬼) "malicious ghost" + Lat. ''draco'' "dragon") is an extinct genus of toothed pterodactyloid pterosaur known from the Early Cretaceous of Liaoning Province, northeast China. According to many recent studies, ''Guidraco'' is a member of the group Anhanguerinae, a subfamily belonging to the larger group Anhangueridae. Discovery ''Guidraco'' is known only from the holotype IVPP V17083, an articulated partial skeleton consisting of a nearly complete skull, lower jaws and a series of four, second to fifth, cervical vertebrae. It was collected at Sihedang near Lingyuan City in the Liaoning Province from the Jiufotang Formation, dating to the Aptian stage of the Early Cretaceous, about 120 million years ago. Etymology ''Guidraco'' was first described and named by Wang Xiaolin, Alexander W.A. Kellner, Jiang Shunxing and Cheng Xin in 2012 and the type species is ''Guidraco venator''. The generic name is derived from Chinese ''gui'' (鬼), "m ...
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Early Cretaceous
The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian-Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining early Aptian Oceanic Anoxic Event 1a (OAE1a) was carbon isotope dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java-Manihiki-Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. The Ontong Java Plateau today covers an area of 1,860,000 km2. In the Indian Ocean another LIP began to form at c. 120 Ma, the Kerguelen P ...
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Mandibular Symphysis
In human anatomy, the facial skeleton of the skull the external surface of the mandible is marked in the median line by a faint ridge, indicating the mandibular symphysis (Latin: ''symphysis menti'') or line of junction where the two lateral halves of the mandible typically fuse at an early period of life (1-2 years). It is not a true symphysis as there is no cartilage between the two sides of the mandible. This ridge divides below and encloses a triangular eminence, the mental protuberance, the base of which is depressed in the center but raised on either side to form the mental tubercle. The lowest (most inferior) end of the mandibular symphysis — the point of the chin — is called the "menton". It serves as the origin for the geniohyoid and the genioglossus muscles. Other animals Solitary mammalian carnivores that rely on a powerful canine bite to subdue their prey have a strong mandibular symphysis, while pack hunters delivering shallow bites have a weaker one. When filter ...
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Anhangueridae
Anhangueridae is a group of pterosaurs within the suborder Pterodactyloidea. They were among the last pterosaurs to possess teeth. A recent study discussing the group considered the Anhangueridae to be typified by a premaxillary crest and a lateral expansion in the distal rostrum. The same study presented a cladistic analysis, for which an "agreement subtree" was calculated. The Anhangueridae was found to be sister taxon to the large crested ''Tropeognathus''. Relationships There are competing theories of ornithocheiromorph phylogeny (evolutionary relationships). Below is cladogram following a topology recovered by Brian Andres, using the most recent iteration of his data set. The cladogram below follows Pêgas ''et al.'' (2019), who recovered Anhangueridae as a much more inclusive group. The analysis found most of the ornithocheirids falling into this family, while ''Ornithocheirus'' itself was recovered as a basal member of Ornithocheirae Ornithocheirae is an extinct clad ...
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