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Lonchodectidae
Lonchodectidae or LonchodraconidaePêgas, R.V., Holgado, B., Leal, M.E.C., 2019. "''Targaryendraco wiedenrothi'' gen. nov. (Pterodactyloidea, Pteranodontoidea, Lanceodontia) and recognition of a new cosmopolitan lineage of Cretaceous toothed pterodactyloids", ''Historical Biology'', 1–15. doi:10.1080/08912963.2019.1690482 is a group of pterosaurs within the clade Pterodactyloidea. It has variously been considered to be within Ctenochasmatoidea, Azhdarchoidea and Pteranodontia.Witton, M.P., Martill, D.M., and Green, M. (2009). "On pterodactyloid diversity in the British Wealden (Lower Cretaceous) and a reappraisal of “Palaeornis” ''cliftii'' Mantell, 1844." ''Cretaceous Research'', 30: 676–686. They are notable for their high, conical tooth sockets and raised alveolar margins. Taxonomic history Lonchodectidae was first named by paleontologist Reginald Walter Hooley in 1914, and was first considered to only contain species of ''Lonchodectes''.Unwin, David M. 2001. "An over ...
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Lonchodectes
''Lonchodectes'' (meaning "lance biter") was a genus of lonchodectid pterosaur from several formations dating to the Turonian (Late Cretaceous) of England, mostly in the area around Kent. The species belonging to it had been assigned to ''Ornithocheirus'' until David Unwin's work of the 1990s and 2000s.Kellner, A.W.A. (2003). Pterosaur phylogeny and comments on the evolutionary history of the group: In: Buffetaut, E., and Mazin, J.-M. (Eds.). ''Evolution and Palaeobiology of Pterosaurs''. Geological Society Special Publication 217:105-137. 1-86239-143-2. Several potential species are known; most are based on scrappy remains, and have gone through several other generic assignments. The genus is part of the complex taxonomy issues surrounding Early Cretaceous pterosaurs from Brazil and England, such as ''Amblydectes'', ''Anhanguera'', ''Coloborhynchus'', and ''Ornithocheirus''. History and species Numerous species have been referred to this genus over time, and only those more ...
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Lonchodraco
''Lonchodraco'' is a genus of lonchodraconid Pterodactyloidea, pterodactyloid pterosaur from the Late Cretaceous of southern England. The genus includes species that were previously assigned to other genera. Discovery and naming In 1846, James Scott Bowerbank named and described some remains found in a chalk pit at Burham near Maidstone in Kent, as a new species of ''Pterodactylus'': ''Pterodactylus giganteus''. The specific name (zoology), specific name means "the gigantic one" in Latin. The same pit generated remains of ''Cimoliopterus, Pterodactylus cuvieri''. In 1848, Bowerbank published a histological study of the bone structure of ''P. giganteus''. At the time, the British Association Code of 1843 allowed to change names if they were inappropriate. In 1850, Richard Owen, considering the species not to have been particularly large, and renamed it into ''Pterodactylus conirostris''; the specific name meaning "cone-snouted", which was based on the conical snout of specimen ...
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Unwindia
''Unwindia'' is a genus of pterodactyloid pterosaur from the Early Cretaceous period (Albian stage) of what is now modern-day Brazil. Discovery and naming The German State Museum of Natural History Karlsruhe obtained a pterosaur fossil from an illegal commercial digger who had found it somewhere near Santana do Cariri in the Araripe Basin. The deposits it was recovered from likely belong to the Santana Formation of northeastern Brazil, which was dated by Martill to the latest Albian stage of the Cretaceous period. The specimen is catalogued as SMNK PAL 6597, and represents a partial skull consisting only the fused premaxillae and maxillae, some teeth and parts of the palate. The frontmost centimeter of the snout tip is missing. Due to the fragility of the specimen, the right side has undergone more preparation. In 2011, the specimen was named and described as the type species ''Unwindia trigonus'' by David Martill. The generic name honours the British pterosaur expert David ...
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Lanceodontia
Ornithocheiromorpha (from Ancient Greek, meaning "bird hand form") is a group of pterosaurs within the suborder Pterodactyloidea. Fossil remains of this group date back from the Early to Late Cretaceous periods (Valanginian to Turonian stages), around 140 to 92.5 million years ago. Ornithocheiromorphs were discovered worldwide except Antarctica, though most genera were recovered in Europe, Asia and South America. They were the most diverse and successful pterosaurs during the Early Cretaceous, but throughout the Late Cretaceous they were replaced by better adapted and more advanced pterosaur species such the pteranodontids and azhdarchoids. The Ornithocheiromorpha was defined in 2014 by Andres and colleagues, and they made Ornithocheiromorpha the most inclusive clade containing ''Ornithocheirus'', but not '' Pteranodon''. Ornithocheiromorphs are considered to be some of the largest animals to have ever flown. Members of this group are also regarded to have some of the lar ...
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Ornithocheiromorpha
Ornithocheiromorpha (from Ancient Greek, meaning "bird hand form") is a group of pterosaurs within the suborder Pterodactyloidea. Fossil remains of this group date back from the Early Cretaceous, Early to Late Cretaceous Period (geology), periods (Valanginian to Turonian stages), around 140 to 92.5 million years ago. Ornithocheiromorphs were discovered worldwide except Antarctica, though most genera were recovered in Europe, Asia and South America. They were the most diverse and successful pterosaurs during the Early Cretaceous, but throughout the Late Cretaceous they were replaced by better adapted and more advanced pterosaur species such the pteranodontids and azhdarchoids. The Ornithocheiromorpha was defined in 2014 by Andres and colleagues, and they made Ornithocheiromorpha the most inclusive clade containing ''Ornithocheirus'', but not ''Pteranodon''. Ornithocheiromorphs are considered to be some of the largest animals to have ever flown. Members of this group are also rega ...
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Lanceodontia
Ornithocheiromorpha (from Ancient Greek, meaning "bird hand form") is a group of pterosaurs within the suborder Pterodactyloidea. Fossil remains of this group date back from the Early to Late Cretaceous periods (Valanginian to Turonian stages), around 140 to 92.5 million years ago. Ornithocheiromorphs were discovered worldwide except Antarctica, though most genera were recovered in Europe, Asia and South America. They were the most diverse and successful pterosaurs during the Early Cretaceous, but throughout the Late Cretaceous they were replaced by better adapted and more advanced pterosaur species such the pteranodontids and azhdarchoids. The Ornithocheiromorpha was defined in 2014 by Andres and colleagues, and they made Ornithocheiromorpha the most inclusive clade containing ''Ornithocheirus'', but not '' Pteranodon''. Ornithocheiromorphs are considered to be some of the largest animals to have ever flown. Members of this group are also regarded to have some of the lar ...
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Boreopteridae
Boreopteridae (meaning "northern wings") is a group of pterodactyloid pterosaurs from the Aptian-age Lower Cretaceous Yixian Formation of Liaoning, China. Classification In 2006, Lü Junchang and colleagues named the clade Boreopteridae for the clade containing the common ancestor of ''Boreopterus'' and ''Feilongus'' and all its descendants, which the authors reclassified as close relatives of the ornithocheirids. (''Feilongus'' had originally been considered a gallodactylid). Many possible boreopterids were subsequently described, one possible example being ''Aetodactylus'', which has been claimed to be similar to ''Boreopterus''. Originally considered close relatives of the ornithocheirids, many of these supposed boreopterids have been found to belong to other groups of the pterodactyloid lineage. In 2012, a phylogenetic analysis by Lü ''et al.'' divided the Boreopteridae into two subfamilies: Boreopterinae, comprising ''Boreopterus'' and ''Zhenyuanopterus'', and Moganopterina ...
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Ikrandraco
''Ikrandraco'' ("Ikran Avatar'' with a crest on the lower jaw] dragon") is a genus of lonchodraconid pterodactyloid pterosaur known from Lower Cretaceous rocks in northeastern China and the Cambridge Greensand of the UK. It is notable for its unusual skull, which features a crest on the lower jaw. Discovery and naming The type species, ''Ikrandraco avatar'' is based on specimen IVPP V18199, a partial skeleton including the skull and jaws, several neck vertebrae, a partial sternal plate, parts of both wings, and part of a foot. A second specimen, IVPP 18406, has also been assigned to ''Ikrandraco''; it consists of a skull and jaws and the first three neck vertebrae. Both specimens come from the Aptian-age Lower Cretaceous Jiufotang Formation of Liaoning, with an estimated date of 120 million years ago. The type species is ''I. avatar'', a second reference to the movie ''Avatar''. It was described in 2014 by Wang Xiaolin and colleagues, but was not properly named a ...
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Zhenyuanopterus
''Zhenyuanopterus'' is a genus of boreopterid pterosaur which is known from Lower Cretaceous (early Aptian stage) Yixian Formation of Liaoning, China. It contains one species, ''Zhenyuanopterus longirostris'', which was first described and named by Lü Junchang. Discovery and naming ''Zhenyuanopterus longirostris'' was named by paleontologist Lü Junchang in 2005. The genus is named after Sun Zhenyuan, who gave Lü Junchang the fossils along with Zi Fan. The specific name ("long-snouted") refers to the long snout of the pterosaur. One specimen is known; a complete skeleton, it was found in the Huangbanjigou locality of the Yixian Formation, and it is catalogued as GLGMV 0001 in the Guilin Longshan Geological Museum in Guangxi, China. A second, larger partial specimen, consisting of the rear half of the body, was found at the same locality and catalogued as XHPM1088 at the Dalian Xinghai Paleontological Museum. In 2014, Teng Fangfang and colleagues described it. Description T ...
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Boreopterus
''Boreopterus'' is a genus of boreopterid pterodactyloid pterosaur from the Barremian-Aptian-age Lower Cretaceous Yixian Formation of Dalian, Liaoning, China. Etymology ''Boreopterus'' was named in 2005 by Lü Junchang and Ji Qiang. The type species is ''Boreopterus cuiae''. The genus name is derived from Greek ''boreios'', "northern" and ''pteron'', "wing". The specific epithet honors Cui Xu. Description ''Boreopterus'' is based on holotype JZMP-04-07-3, a nearly complete but crushed skeleton and skull. The skull is 235 millimeters long (9.25 inches), low and elongated with a rounded tip. Its wingspan is estimated to have been around 1.45 meters (4.76 feet). Its teeth, especially the anterior nine pairs, are quite large, forming a mesh of sharp teeth at the front of the mouth; the third and fourth teeth from the front are the largest. There are at least 27 teeth in each side of both the upper and lower jaws, which is a large amount. Classification Lü and Ji ...
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Yixianopterus
''Yixianopterus'' is a pterodactyloid pterosaur genus from the Barremian-Aptian-age Lower Cretaceous Yixian Formation of Liaoning, China. It is known from a single specimen, holotype JZMP-V-12, housed at the Benxi Geological Museum. This specimen was doctored before acquisition, with much of the skull being fabricated. Classification Lü ''et al.'' (2006) assigned ''Yixianopterus'' to the lophocratian family Lonchodectidae on the basis of dental characters, classifying it as the first Asian representative of Lonchodectidae. Martill (2011) considered it potentially related to his new taxon ''Unwindia'', and Witton (2013) assigned it to the family Ornithocheiridae.Witton, Mark P. (2013). ''Pterosaurs: Natural History, Evolution, Anatomy.'' Princeton, NJ: Princeton University Press. A reappraisal of the holotype specimen and phylogenetic analysis by Jiang ''et al.'' (2020) recovered ''Yixianopterus'' as a basal member of the clade Istiodactyliformes Ornithocheiromorpha (from An ...
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Serradraco
''Serradraco'' is a genus of Early Cretaceous pterodactyloid pterosaur from the Valanginian aged Tunbridge Wells Sand Formation in England. Named by Rigal ''et al.'' in 2018 with the description of a second specimen, it contains a single species, ''S. sagittirostris'', which was formerly considered a species of ''Lonchodectes'', ''L. sagittirostris''. In 2020, Averianov suggested it did not belong in Lonchodectidae. The second specimen, BEXHM 2015.18, consists of a small fragment of jaw with three teeth; six complete caudal vertebrae fused together and two fragmentary caudal vertebrae in articulation; a distal left ulna; a right proximal syncarpal; portions of a minimum of three phalanges and two indeterminate elements. Discovery and naming In 1874, Richard Owen named a pair of lower jaws from the collection of Samuel Husband Beckles, found at St Leonards-on-Sea in Sussex, as a new species of ''Pterodactylus'': ''Pterodactylus sagittirostris''. The specific name means "arrowhead ...
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