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Zoospores Types
A zoospore is a motile asexual spore that uses a flagellum for locomotion. Also called a swarm spore, these spores are created by some protists, bacteria, and fungi to propagate themselves. Diversity Flagella types Zoospores may possess one or more distinct types of flagella - tinsel or "decorated", and whiplash, in various combinations. *Tinsellated (straminipilous) flagella have lateral filaments known as mastigonemes perpendicular to their main axis, which allow for more surface area, and disturbance of the medium, giving them the property of a rudder, that is, used for steering. *Whiplash flagella are straight, to power the zoospore through its medium. Also, the "default" zoospore only has the propelling, whiplash flagella. Both tinsel and whiplash flagella beat in a sinusoidal wave pattern, but when both are present, the tinsel beats in the opposite direction of the whiplash, to give two axes of control of motility. Morphological types In eukaryotes, the four main typ ...
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Chytridiomycota
Chytridiomycota are a division of zoosporic organisms in the kingdom Fungi, informally known as chytrids. The name is derived from the Ancient Greek ('), meaning "little pot", describing the structure containing unreleased zoöspores. Chytrids are one of the early diverging fungal lineages, and their membership in kingdom Fungi is demonstrated with chitin cell walls, a posterior whiplash flagellum, absorptive nutrition, use of glycogen as an energy storage compound, and synthesis of lysine by the -amino adipic acid (AAA) pathway. Chytrids are saprobic, degrading refractory materials such as chitin and keratin, and sometimes act as parasites. There has been a significant increase in the research of chytrids since the discovery of '' Batrachochytrium dendrobatidis'', the causal agent of chytridiomycosis. Classification Species of Chytridiomycota have traditionally been delineated and classified based on development, morphology, substrate, and method of zoöspore discha ...
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Gametangium
A gametangium (plural: gametangia) is an organ or cell in which gametes are produced that is found in many multicellular protists, algae, fungi, and the gametophytes of plants. In contrast to gametogenesis in animals, a gametangium is a haploid structure and formation of gametes does not involve meiosis. Types of gametangia Depending on the type of gamete produced in a gametangium, several types can be distinguished. Female Female gametangia are most commonly called archegonia. They produce egg cells and are the sites for fertilization. Archegonia are common in algae and primitive plants as well as gymnosperms. In flowering plants, they are replaced by the embryo sac inside the ovule. Male The male gametangia are most commonly called antheridia. They produce sperm cells that they release for fertilization. Antheridia producing non-motile sperm (spermatia) are called spermatangia. Some antheridia do not release their sperm. For example, the oomycete antheridium is a syncytiu ...
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Fern
A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except the lycopods, and differ from mosses and other bryophytes by being vascular, i.e., having specialized tissues that conduct water and nutrients and in having life cycles in which the branched sporophyte is the dominant phase. Ferns have complex leaves called megaphylls, that are more complex than the microphylls of clubmosses. Most ferns are leptosporangiate ferns. They produce coiled fiddleheads that uncoil and expand into fronds. The group includes about 10,560 known extant species. Ferns are defined here in the broad sense, being all of the Polypodiopsida, comprising both the leptosporangiate ( Polypodiidae) and eusporangiate ferns, the latter group including horsetails, whisk ferns, marattioid ferns, and ophioglossoid ...
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Angiosperm
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants that produce their seeds enclosed within a fruit. They are by far the most diverse group of land plants with 64 orders, 416 families, approximately 13,000 known genera and 300,000 known species. Angiosperms were formerly called Magnoliophyta (). Like gymnosperms, angiosperms are seed-producing plants. They are distinguished from gymnosperms by characteristics including flowers, endosperm within their seeds, and the production of fruits that contain the seeds. The ancestors of flowering plants diverged from the common ancestor of all living gymnosperms before the end of the Carboniferous, over 300 million years ago. The closest fossil relatives of flowering plants are uncertain and contentious. The earliest angiosperm fossils are in ...
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Sporangium
A sporangium (; from Late Latin, ) is an enclosure in which spores are formed. It can be composed of a single cell or can be multicellular. Virtually all plants, fungi, and many other lineages form sporangia at some point in their life cycle. Sporangia can produce spores by mitosis, but in nearly all land plants and many fungi, sporangia are the site of meiosis and produce genetically distinct haploid spores. Fungi In some phyla of fungi, the sporangium plays a role in asexual reproduction, and may play an indirect role in sexual reproduction. The sporangium forms on the sporangiophore and contains haploid nuclei and cytoplasm. Spores are formed in the sporangiophore by encasing each haploid nucleus and cytoplasm in a tough outer membrane. During asexual reproduction, these spores are dispersed via wind and germinate into haploid hyphae. Although sexual reproduction in fungi varies between phyla, for some fungi the sporangium plays an indirect role in sexual reprod ...
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Heterokont
Heterokonts are a group of protists (formally referred to as Heterokonta, Heterokontae or Heterokontophyta). The group is a major line of eukaryotes. Most are algae, ranging from the giant multicellular kelp to the unicellular diatoms, which are a primary component of plankton. Other notable members of the Stramenopiles include the (generally) parasitic oomycetes, including '' Phytophthora'', which caused the Great Famine of Ireland, and '' Pythium'', which causes seed rot and damping off. The name "heterokont" refers to the type of motile life cycle stage, in which the flagellated cells possess two differently arranged flagella (see zoospore). History In 1899, Alexander Luther created the term "Heterokontae" for some algae with unequal flagella, today called Xanthophyceae. Later, some authors (e.g., Copeland, 1956) included other groups in Heterokonta, expanding the name's sense. The term continues to be applied in different ways, leading to Heterokontophyta being appl ...
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Oomycota
Oomycota forms a distinct phylogenetic lineage of fungus-like eukaryotic microorganisms, called oomycetes (). They are filamentous and heterotrophic, and can reproduce both sexually and asexually. Sexual reproduction of an oospore is the result of contact between hyphae of male antheridia and female oogonia; these spores can overwinter and are known as resting spores. Asexual reproduction involves the formation of chlamydospores and sporangia, producing motile zoospores. Oomycetes occupy both saprophytic and pathogenic lifestyles, and include some of the most notorious pathogens of plants, causing devastating diseases such as late blight of potato and sudden oak death. One oomycete, the mycoparasite ''Pythium oligandrum'', is used for biocontrol, attacking plant pathogenic fungi. The oomycetes are also often referred to as water molds (or water moulds), although the water-preferring nature which led to that name is not true of most species, which are terrestrial pathogens. ...
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Hyphochytriomycetes
Hyphochytrids are eukaryotic organisms in the group of Stramenopiles (Heterokonta). Characteristics They are distinguished by an anterior tinsel flagellum on their zoospores. Also they have a rhizoidal or hypha-like vegetative system (hence the prefix "Hypho-"). Classification This group may be put alternatively at the phylum, class, subclass or order level, being referred to as Hyphochytriomycota, Hyphochytriomycetes (or Hyphochytrea), Hyphochytriomycetidae (or Hyphochytridae) and Hyphochytriales, respectively. The variants Hyphochytri''di''omycota and Hyphochytri''di''omycetes are also sometimes used, presumably by analogy to the Chytridiomycetes, or due to the perpetuation of a typographical error. However, the stem is Hyphochytri- (from '' Hyphochytrium'') and not Hyphochytridi- (from ''Chytridium''). In the past the classes Hyphochytridiomycetes, Oomycetes and Chytridiomycetes were grouped together in the now obsolete taxon Mastigomycotina as fungi with flagellate spores ...
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Plasmodiophoromycota
The plasmodiophores (also known as plasmophorids or plasmodiophorids) are a group of obligate endoparasitic protists belonging to the subphylum Endomyxa in Cercozoa. Taxonomically, they are united under a single family Plasmodiophoridae, order Plasmodiophorida, sister to the phagomyxids. Ecology and pathology Plasmodiophores are pathogenic for a wide range of organisms, but mainly green plants. The more commonly recognized are agents of plant diseases such as clubroot, powdery scab and crook root of watercress, or vectors for viruses that infect beets, peanut, monocots and potatoes, such as the potato mop-top virus or the beet necrotic yellow vein virus. Taxonomy History The plasmodiophores have historically been regarded as Fungi. The first description of plasmodiophores as a taxonomic group was in 1885 by Zopf, who united two genera '' Plasmodiophora'' and '' Tetramyxa'' in a common family “Plasmodiophoreæ”, inside the group “Monadineæ”, as part of the division ...
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Myxogastria
Myxogastria/Myxogastrea (myxogastrids, ICZN) or Myxomycetes ( ICN), is a class of slime molds that contains 5 orders, 14  families, 62 genera, and 888 species. They are colloquially known as the ''plasmodial'' or ''acellular'' slime moulds. All species pass through several, very different morphologic phases, such as microscopic individual cells, slimy amorphous organisms visible with the naked eye and conspicuously shaped fruit bodies. Although they are monocellular, they can reach immense widths and weights: in extreme cases they can be up to across and weigh up to . The class Myxogastria is distributed worldwide, but it is more common in temperate regions where it has a higher biodiversity than in polar regions, the subtropics or tropics. They are mainly found in open forests, but also in extreme regions such as deserts, under snow blankets or underwater. They also occur on the bark of trees, sometimes high in the canopy. These are known as corticolous ...
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Neocallimastigales
Neocallimastigomycota is a phylum containing anaerobic fungi, which are symbionts found in the digestive tracts of larger herbivores. Anaerobic fungi were originally placed within phylum Chytridiomycota, within Order Neocallimastigales but later raised to phylum level, a decision upheld by later phylogenetic reconstructions. It encompasses only one family. Discovery The fungi in Neocallimastigomycota were first recognised as fungi by Orpin in 1975, based on motile cells present in the rumen of sheep. Their zoospores had been observed much earlier but were believed to be flagellate protists, but Orpin demonstrated that they possessed a chitin cell wall. It has since been shown that they are fungi related to the core chytrids. Prior to this, the microbial population of the rumen was believed to consist only of bacteria and protozoa. Since their discovery they have been isolated from the digestive tracts of over 50 herbivores, including ruminant and non-ruminant (hindgut-fermenting ...
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