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Waeringopteroidea
Waeringopteridae is a family of eurypterids, an extinct group of aquatic arthropods. The Waeringopteridae is the only family classified as part of the superfamily Waeringopteroidea, which in turn is classified within the infraorder Diploperculata in the suborder Eurypterina. The earliest known member of the group, '' Orcanopterus'', has been recovered from deposits of Katian (Late Ordovician) age and the latest known surviving member, ''Grossopterus'', has been recovered from deposits of Siegenian (Early Devonian) age. The name Waeringopteridae is derived from the type genus '' Waeringopterus'', which is named in honor of eurypterid researcher Erik N. Kjellesvig-Waering. Waeringopterid fossils are very rare and the family only contains five known species classified in three different genera (''Waeringopterus'', ''Orcanopterus'' and ''Grossopterus''). Four of these species are from the ancient continent of Laurentia, with the only species known from outside of it being a Devonian ...
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Eurypterina
Eurypterina is one of two suborders of eurypterids, an extinct group of chelicerate arthropods commonly known as "sea scorpions". Eurypterine eurypterids are sometimes informally known as "swimming eurypterids". They are known from fossil deposits worldwide, though primarily in North America and Europe. Seventy-five percent of eurypterid species are eurypterines; this represents 99% of specimens. The superfamily Pterygotioidea is the most species-rich clade, with 56 species, followed by the Adelophthalmoidea with 43 species; as sister taxa, they comprise the most derived eurypterines. Pterygotioidea includes the pterygotids, which are the only eurypterids known to have a cosmopolitan distribution. Though more numerous both in specimens and taxa, the eurypterines have the shorter temporal range of the two eurypterid suborders. They first appeared around the same time as the Stylonurina in the Middle Ordovician. The suborder faced a slow extinction during the Middle and Late Devo ...
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Adelophthalmidae
Adelophthalmidae (the name deriving from the type genus ''Adelophthalmus'', meaning "no obvious eyes") is a family of eurypterids, an extinct group of aquatic arthropods. Adelophthalmidae is the only family classified as part of the superfamily Adelophthalmoidea, which in turn is classified within the infraorder Diploperculata in the suborder Eurypterina. Adelophthalmid eurypterids were small and swimming eurypterids that appeared in the Silurian period. With the earliest known members of the group, '' Nanahughmilleria prominens'' and '' Parahughmilleria maria'', being known from deposits of Early Silurian (possibly the Llandovery epoch) age and the last members, belonging to the long-lasting and widespread genus ''Adelophthalmus'', going extinct in the Early Permian, the Adelophthalmidae is the longest lasting single family of eurypterids. The survival of the group, and of swimming eurypterids (the suborder Eurypterina) beyond the Late Devonian is entirely due to the survival, a ...
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Diploperculata
Diploperculata is an infraorder of eurypterids, an extinct group of aquatic arthropods commonly known as "sea scorpions". The name, derived from Greek διπλόω ("double") and ''operculum'', refers to the distinguishing feature that unites the superfamilies included in the group, that the genital operculum is made up of two fused segments. The group contains the three eurypterine superfamilies Carcinosomatoidea, Adelophthalmoidea and Pterygotioidea alongside the as of yet undescribed " waeringopteroid" clade. The diploperculate eurypterine clade had prior to its description been noted as a monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ... group of derived eurypterines that form a sister-taxon to the superfamily Eurypteroidea. See also * List of eurypterid gen ...
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Adelophthalmoidea
Adelophthalmidae (the name deriving from the type genus ''Adelophthalmus'', meaning "no obvious eyes") is a family of eurypterids, an extinct group of aquatic arthropods. Adelophthalmidae is the only family classified as part of the superfamily Adelophthalmoidea, which in turn is classified within the infraorder Diploperculata in the suborder Eurypterina. Adelophthalmid eurypterids were small and swimming eurypterids that appeared in the Silurian period. With the earliest known members of the group, '' Nanahughmilleria prominens'' and '' Parahughmilleria maria'', being known from deposits of Early Silurian (possibly the Llandovery epoch) age and the last members, belonging to the long-lasting and widespread genus ''Adelophthalmus'', going extinct in the Cisuralian, Early Permian, the Adelophthalmidae is the longest lasting single family of eurypterids. The survival of the group, and of swimming eurypterids (the suborder Eurypterina) beyond the Late Devonian is entirely due to the ...
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Eurypterid
Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the Order (biology), order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 Myr, million years ago. The group is likely to have appeared first either during the Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic Chelicerata, chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian, from which the majority of eurypterid species have been described. The Silurian genus ''Eurypterus'' accounts for more than 90% of all known eurypterid specimens. Though the group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event. They declined in numbers and diversity until becoming extinct during the Permian–Tri ...
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Pterygotid
Pterygotidae (the name deriving from the type genus ''Pterygotus'', meaning "winged one") is a family of eurypterids, an extinct group of aquatic arthropods. They were members of the superfamily Pterygotioidea. Pterygotids were the largest known arthropods to have ever lived with some members of the family, such as ''Jaekelopterus'' and '' Acutiramus'', exceeding 2 metres (6.6 ft) in length. Their fossilized remains have been recovered in deposits ranging in age from 428 to 372 million years old (Late Silurian to Late Devonian). One of the most successful groups of eurypterids, the pterygotids were the only eurypterid family to achieve a truly worldwide distribution. Several evolutionary innovations made the pterygotids unique among the eurypterids, with large and flattened telsons (the posteriormost division of the body) likely used as rudders to provide additional agility and enlarged chelicerae (frontal appendages) with claws. These claws were robust and possessed teeth ...
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Pterygotioidea
Pterygotioidea (the name deriving from the type genus ''Pterygotus'', meaning "winged one") is a superfamily of eurypterids, an extinct group of aquatic arthropods. Pterygotioids were the most derived members of the infraorder Diploperculata and the sister group of the adelophthalmoid eurypterids. The group includes the basal and small hughmilleriids, the larger and specialized slimonids and the famous pterygotids which were equipped with robust and powerful cheliceral claws. Though the more primitive hughmilleriids were small, '' Hughmilleria wangi'' being the smallest of all pterygotioids at just in length, later members of the group, particularly in the Pterygotidae, would become the largest known arthropods to ever exist with several genera surpassing in length. Among all currently recognized eurypterid clades, the Pterygotioidea is the most diverse, containing over 50 species in 10 genera. With the number of recognized eurypterid species being around 250, pterygotio ...
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Biology Letters
''Biology Letters'' is a peer-reviewed, biological, scientific journal published by the Royal Society. It focuses on the rapid publication of short high quality research articles, reviews and opinion pieces across the biological sciences. ''Biology Letters'' has an average turnaround time of twenty four days from submission to a first decision. The editor-in-chief is Professor David Beerling FRS (University of Sheffield) who is supported by an international Editorial Board of practising scientists. Contents and themes As well as the conventional, short research articles, ''Biology Letters'' has recently published Special Features and Mini Series. While Special Features are a collection of up to 20 articles on a specific theme and published across multiple issues, Mini Series include up to six articles that are published in one issue. Examples of topics in these formats include ocean acidification, fossils, extinction, enhanced rock weathering and the evolutionary ecology of sp ...
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Carcinosomatid
Carcinosomatidae (the name deriving from the type genus ''Carcinosoma'', meaning "crab body")Meaning osomaat ''www.dictionary.com''. Retrieved 7 September 2018. is a family of eurypterids, an extinct group of aquatic arthropods. They were members of the superfamily Carcinosomatoidea, also named after ''Carcinosoma''. Fossils of carcinosomatids have been found in North America, Europe and Asia, the family possibly having achieved a worldwide distribution, and range in age from the Late Ordovician to the Early Devonian. They were among the most marine eurypterids, known almost entirely from marine environments. Carcinosomatids varied considerably in size, from species only a few centimetres in length to some of the largest known arthropods. The largest carcinosomatid species, ''Carcinosoma punctatum'', reached lengths of at least and rivalled the largest eurypterid of all, ''Jaekelopterus'', in size. Morphologically, carcinosomatids were highly distinct from other eurypterids, know ...
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Cuticle
A cuticle (), or cuticula, is any of a variety of tough but flexible, non-mineral outer coverings of an organism, or parts of an organism, that provide protection. Various types of "cuticle" are non- homologous, differing in their origin, structure, function, and chemical composition. Human anatomy In human anatomy, "cuticle" can refer to several structures, but it is used in general parlance, and even by medical professionals, to refer to the thickened layer of skin surrounding fingernails and toenails (the eponychium), and to refer to the superficial layer of overlapping cells covering the hair shaft ( cuticula pili), consisting of dead cells, that locks the hair into its follicle. It can also be used as a synonym for the epidermis, the outer layer of skin. Cuticle of invertebrates In zoology, the invertebrate cuticle or cuticula is a multi-layered structure outside the epidermis of many invertebrates, notably roundworms and arthropods, in which it forms an exoskeleton ...
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Carapace
A carapace is a Dorsum (biology), dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the underside is called the plastron. Crustaceans In crustaceans, the carapace functions as a protective cover over the cephalothorax (i.e., the fused head and thorax, as distinct from the abdomen behind). Where it projects forward beyond the eyes, this projection is called a rostrum (anatomy), rostrum. The carapace is Calcification, calcified to varying degrees in different crustaceans. Zooplankton within the phylum Crustacea also have a carapace. These include Cladocera, ostracods, and Isopoda, isopods, but isopods only have a developed "cephalic shield" carapace covering the head. Arachnids In arachnids, the carapace is formed by the fusion of prosomal tergites into a single Plate (animal anatomy), plate which carries the e ...
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Telson
The telson () is the posterior-most division of the body of an arthropod. Depending on the definition, the telson is either considered to be the final segment of the arthropod body, or an additional division that is not a true segment on account of not arising in the embryo from teloblast areas as other segments. It never carries any appendages, but a forked "tail" called the caudal furca may be present. The shape and composition of the telson differs between arthropod groups. Crustaceans In lobsters, shrimp and other decapods, the telson, along with the uropods, forms the tail fan. This is used as a paddle in the caridoid escape reaction ("lobstering"), whereby an alarmed animal rapidly flexes its tail, causing it to dart backwards. Krill can reach speeds of over 60 cm per second by this means. The trigger time to optical stimulus is, in spite of the low temperatures, only 55 milliseconds. In the Isopoda and Tanaidacea (superorder Peracarida), the last abdominal b ...
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