Villarroelia
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Villarroelia
''Villarroelia'' is an extinct genus of proterotheriid from the Middle Miocene of Colombia. Etymology The genus name, ''Villarroelia'' refers to the paleontologist Carlos Villarroel, for his contributions to the understanding of prehistoric South American mammals, while the specific name, ''V. toyotoi'' refers to the indigenous chief Toyotó, whose tribe was present in the area of the Villavieja River, in the present-day Huila Department in Colombia, where its fossils were discovered in sediments corresponding to the La Victoria Formation, belonging to the Honda Group of the Middle Miocene.Cifelli, R. L. and Guerrero, J. (1997). Litopterns. In: ''Vertebrate Paleontology in the Neotropics: The Miocene Fauna of La Venta, Colombia'' (R. Kay, R. H. Madden, R. L. Cifelli, and J. J. Flynn, eds.). Smithsonian Institution Press, Washington and London, 289-302. Description This animal is known from the holotype specimen IGM 250965, a partial skull lacking the anterior rostrum, althoug ...
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Laventan
The Laventan ( es, Laventense) age is a period of geologic time (13.8 to 11.8 Ma) within the Middle Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Colloncuran and precedes the Mayoan age.Madden et al., 1997 Etymology The age is named after the Miocene Lagerstätte La Venta, where a rich biodiversity from the Middle Miocene has been recovered from the Honda Group. Formations Fossil content Correlations The Laventan (13.8 to 11.8 Ma) correlates with: * NALMA ** latest Barstovian (15.97-13.65 Ma)Barstovian
at Fossilworks.org
** early

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Honda Group, Colombia
The Honda Group ( es, Grupo Honda, Tsh, Ngh) is a geological group of the Upper and Middle Magdalena Basins and the adjacent Central and Eastern Ranges of the Colombian Andes. The group, in older literature also defined as formation, is in its present-day type section in the Tatacoa Desert in the department of Huila subdivided into two main formations; La Victoria and Villavieja. The group was originally defined in and named after Honda, Tolima, but has been redefined based on the many fossil finds in the Tatacoa Desert, to the south. In the original type section of its occurrence, the thick group is subdivided into three formations, from old to young; Cambrás, San Antonio and Los Limones. The group dates to the Neogene period; in its broadest definition from the Late Oligocene to Late Miocene, and in the redefined type section restricted to the Laventan age of the South American Land Mammal Ages (SALMA), equivalent to the Middle Miocene Serravallian epoch. The Honda Gr ...
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Proterotheriidae
Proterotheriidae is an extinction, extinct family of fossil ungulates from the Cenozoic era that displays toe reduction. Despite resembling primitive, small horses, they were only distantly related to them, and instead belonged to the native South American ungulate order Litopterna. Description Two subfamilies and 18 genera of Proterotheriidae are known. All forms were small or medium-sized. Typical is a reduction of the number of toes and brachydont or mesodont teeth. The family is recorded since the late Palaeocene. Various fossils are known from many parts of the South American continent. The diversity decreased in the Miocene to Pliocene and it has been assumed for a long time that they entirely disappeared in the late Pliocene. However, fossils found in Argentina, Brazil and Uruguay show that one member of the group, ''Neolicaphrium recens'' survived into the Late Lujanian, Pleistocene. Better known genera of the family include ''Diadiaphorus'' and ''Thoatherium'' from th ...
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Proterotheriids
Proterotheriidae is an extinct family of fossil ungulates from the Cenozoic era that displays toe reduction. Despite resembling primitive, small horses, they were only distantly related to them, and instead belonged to the native South American ungulate order Litopterna. Description Two subfamilies and 18 genera of Proterotheriidae are known. All forms were small or medium-sized. Typical is a reduction of the number of toes and brachydont or mesodont teeth. The family is recorded since the late Palaeocene. Various fossils are known from many parts of the South American continent. The diversity decreased in the Miocene to Pliocene and it has been assumed for a long time that they entirely disappeared in the late Pliocene. However, fossils found in Argentina, Brazil and Uruguay show that one member of the group, '' Neolicaphrium recens'' survived into the Late Pleistocene. Better known genera of the family include '' Diadiaphorus'' and '' Thoatherium'' from the Miocene. Ta ...
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Proterotherium
''Proterotherium'' (meaning "first beast") is an extinct genus of litoptern mammal of the family Proterotheriidae that lived during the Late Miocene of Argentina and Chile. Fossils of this genus have been found in the Ituzaingó Formation of Argentina, and the Galera Formation of Chile. Classification The genus ''Proterotherium'' was first described by Florentino Ameghino in 1883, on the basis of fossil remains found in Upper Miocene deposits in Patagonia. The type species is ''Proterotherium cervioides''. Numerous other species have been ascribed to this genus, many of which come from lower-middle Miocene deposits, such as ''P. americanum'', ''P. australe'', ''P. brachygnathum'', ''P. cavum'', ''P. cingulatum'', ''P. curtidens'', ''P. dichotomum'' , ''P. divortium'', ''P. gradatum'', ''P. intermedium'', ''P. karaikense'', ''P. mixtum'', ''P. nitens'', ''P. perpolitum'', ''P. politum'', ''P. main'', and ''P. pyramidatum''. However, many of these species are now considered to be ...
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Middle Miocene
The Middle Miocene is a sub-epoch of the Miocene Epoch made up of two stages: the Langhian and Serravallian stages. The Middle Miocene is preceded by the Early Miocene. The sub-epoch lasted from 15.97 ± 0.05 Ma to 11.608 ± 0.005 Ma (million years ago). During this period, a sharp drop in global temperatures took place. This event is known as the Middle Miocene Climate Transition. For the purpose of establishing European Land Mammal Ages The European Land Mammal Mega Zones (abbreviation: ELMMZ, more commonly known as European land mammal ages or ELMA) are zones in rock layers that have a specific assemblage of fossils (biozones) based on occurrences of fossil assemblages of Europe ... this sub-epoch is equivalent to the Astaracian age. External links GeoWhen Database - Middle Miocene .02 02 * * {{geochronology-stub ...
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Calcaneus
In humans and many other primates, the calcaneus (; from the Latin ''calcaneus'' or ''calcaneum'', meaning heel) or heel bone is a bone of the tarsus of the foot which constitutes the heel. In some other animals, it is the point of the hock. Structure In humans, the calcaneus is the largest of the tarsal bones and the largest bone of the foot. Its long axis is pointed forwards and laterally. The talus bone, calcaneus, and navicular bone are considered the proximal row of tarsal bones. In the calcaneus, several important structures can be distinguished:Platzer (2004), p 216 There is a large calcaneal tuberosity located posteriorly on plantar surface with medial and lateral tubercles on its surface. Besides, there is another peroneal tubecle on its lateral surface. On its lower edge on either side are its lateral and medial processes (serving as the origins of the abductor hallucis and abductor digiti minimi). The Achilles tendon is inserted into a roughened area on its superio ...
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Neogene Colombia
The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period Mya. The Neogene is sub-divided into two epochs, the earlier Miocene and the later Pliocene. Some geologists assert that the Neogene cannot be clearly delineated from the modern geological period, the Quaternary. The term "Neogene" was coined in 1853 by the Austrian palaeontologist Moritz Hörnes (1815–1868). During this period, mammals and birds continued to evolve into modern forms, while other groups of life remained relatively unchanged. The first humans (''Homo habilis'') appeared in Africa near the end of the period. Some continental movements took place, the most significant event being the connection of North and South America at the Isthmus of Panama, late in the Pliocene. This cut off the warm ocean currents from the Pacific to the ...
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Miocene Mammals Of South America
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the late ...
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Prehistoric Placental Genera
Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of symbols, marks, and images appears very early among humans, but the earliest known writing systems appeared 5000 years ago. It took thousands of years for writing systems to be widely adopted, with writing spreading to almost all cultures by the 19th century. The end of prehistory therefore came at very different times in different places, and the term is less often used in discussing societies where prehistory ended relatively recently. In the early Bronze Age, Sumer in Mesopotamia, the Indus Valley Civilisation, and ancient Egypt were the first civilizations to develop their own scripts and to keep historical records, with their neighbors following. Most other civilizations reached the end of prehistory during the following Iron Age. T ...
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Phylogenetic
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does ...
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Anisolophus
''Anisolophus'' is an extinct genus of proterotheriid from the Early to Middle Miocene of Argentina. The genus was named by Burmeister in 1885 to accommodate the species ''Anchitherium australe'', which they had named earlier in 1879. Soria then referred the species ''Licaphrium floweri'' and ''Anisolophus minisculus'' to the genus, making ''Licaphrium'', named in 1887 by Florentino Ameghino, a junior synonym of the genus. Both ''A. australis'' and ''A. floweri'' are known from the Santacrucian age Santa Cruz Formation, while ''A. minisculus'' is known from the Collón Curá Formation. ''Anisolophus'' is considered the senior synonym of the genus ''Licaphrium'', which was named in 1887 by Ameghino for the species ''L. floweri'', now ''A. floweri''. Many other species of ''Licaphrium'' were named, many of which are considered synonyms of ''A. floweri'', '' Tetramerorhinus'', ''Neobrachytherium'', or '' Lophogonodon'', as well as the dubious Doubt is a mental state in which the ...
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