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Vytshegdosuchus
''Vytshegdosuchus'' is an extinct genus of paracrocodylomorph archosaur known from the Early Triassic (latest Olenekian stage) Yarenskian Gorizont of the Komi Republic of the European section of Russia. It contains a single species, ''Vytshegdosuchus zheshartensis''. ''Vytshegdosuchus'' was named by Andreii Sennikov in 1988. Discovery ''Vytshegdosuchus'' is known from the holotype PIN 3361/134, a right ilium, and possibly from the paratype PIN 3361/127, a partial femur. These specimens were collected from the a fossil assemblage called the ''Parotosuchus'' fauna of the Upper Yarenskian Gorizont, in the Zheshart locality along the Vychegda River in Aikino district of the Komi Republic. A pterygoid and postcranial material from the Gam locality, a cervical vertebra (PIN 3369/139) from another locality and postcrania from the Mezhog locality were also assigned to ''Vytshegdosuchus'' by Sennikov (1999). Nesbitt (2009) argued that the specimens referred to ''Vytshegd ...
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Xilousuchus
''Xilousuchus'' is an extinct genus of poposauroid from lower Triassic (Olenekian stage) deposits of Fugu County of northeastern Shanxi Province, China. It is known from the holotype, IVPP V 6026, a single well-preserved partial skeleton including the skull. It was found from the Heshanggou Formation of the Ordos Basin, Hazhen commune. It was first named by Xiao-Chun Wu in 1981 and the type species is ''Xilousuchus sapingensis''. Wu (1981) referred ''Xilousuchus'' to the Proterosuchia. Gower and Sennikov (1996) found it to be an erythrosuchian based strictly on the braincase. A more detailed re-description of the genus was provided by Nesbitt ''et al.'' (2010) and found poposauroid affinities. In his massive revision of archosaurs which included a large cladistic analysis, Sterling J. Nesbitt (2011) found ''Xilousuchus'' to be a poposauroid which is most closely related to ''Arizonasaurus''. ''Xilousuchus'' is the oldest archosaur to date, although ''Ctenosauriscus'' ...
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Paracrocodylomorph
Paracrocodylomorpha is a clade of pseudosuchian archosaurs. The clade includes the diverse and unusual group Poposauroidea as well as the generally carnivorous and quadrupedal members of Loricata, including modern crocodylians. Paracrocodylomorpha was named by paleontologist J. Michael Parrish in 1993, although the group is now considered to encompass more reptiles than his original definition intended. The most recent definition of Paracrocodylomorpha, as defined by Sterling Nesbitt in 2011, is "the least inclusive clade containing ''Poposaurus'' and ''Crocodylus niloticus'' (the Nile crocodile). Most groups of paracrocodylomorphs became extinct at the end of the Triassic period, with the exception of the crocodylomorphs, from which crocodylians such as crocodiles and alligators evolved in the latter part of the Mesozoic. History and definition Parrish (1993) defined Paracrocodylomorpha as the last common ancestor of "Poposauridae" and Crocodylomorpha and all of its descenda ...
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Early Triassic
The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between Ma and Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series, which is a unit in chronostratigraphy. The Early Triassic is the oldest epoch of the Mesozoic Era. It is preceded by the Lopingian Epoch (late Permian, Paleozoic Era) and followed by the Middle Triassic Epoch. The Early Triassic is divided into the Induan and Olenekian ages. The Induan is subdivided into the Griesbachian and Dienerian subages and the Olenekian is subdivided into the Smithian and Spathian subages. The Lower Triassic series is coeval with the Scythian Stage, which is today not included in the official timescales but can be found in older literature. In Europe, most of the Lower Triassic is composed of Buntsandstein, a lithostratigraphic unit of continental red beds. The Early Triassic and partly also the Middle Triassic span the in ...
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Vychegda River
, image = VytchegdaRiver.jpg , image_size = , image_caption = The lower course of the Vychegda , source1_location = , mouth = Northern Dvina , mouth_coordinates = , progression = , subdivision_type1 = Country , subdivision_name1 = Russia , length = , source1_elevation = , mouth_elevation = , discharge1_avg = , basin_size = , tributaries_left = North Keltma, Sysola, Viled , tributaries_right = Vishera, Vym, Yarenga, Berdyshevka The Vychegda (, ) is a river in the European part of Russia, tributary to the Northern Dvina. Its length is about . Its source is approximately west of the northern Ural Mountains. It flows roughly in western direction, through Komi Republic and Arkhangelsk Oblast. The largest city along the Vychegda is Syktyvkar, the capital of Komi Republic. The Viled, the Yarenga, and the Vym are among its main tributaries. The Vychegda flows into ...
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Poposauroid
Poposauroidea is a clade of advanced pseudosuchians (archosaurs closer to crocodilians than to dinosaurs). It includes poposaurids, shuvosaurids, ctenosauriscids, and other unusual pseudosuchians such as ''Qianosuchus'' and ''Lotosaurus''. However, it excludes most large predatory quadrupedal "rauisuchians" such as rauisuchids and "prestosuchids". Those reptiles are now allied with crocodylomorphs (crocodile ancestors) in a clade known as Loricata, which is the sister taxon to the poposauroids in the clade Paracrocodylomorpha. Although it was first formally defined in 2007, the name "Poposauroidea" has been used for many years. The group has been referred to as Poposauridae by some authors, although this name is often used more narrowly to refer to the family that includes ''Poposaurus'' and its close relatives. Poposauroids went extinct at the end of the Triassic period along with other non-crocodylomorph pseudosuchians. However, they were among the most diverse and long-lasting ...
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Fugusuchus
''Fugusuchus'' is an extinct genus of archosauriform, probably the basal-most member of the family Erythrosuchidae. The genus is known from a single fossil from the middle Early Triassic Heshanggou Formation in Shanxi, China. The partial skeleton consists of an incomplete skull, parts of the right forelimb, and an intercentrum. The skeleton, known as GMB V 313, is currently in the Geological Museum of China in Beijing. ''Fugusuchus'' was a medium-sized archosauriform. It has a long and relatively low skull, unlike the higher more pointed skulls of related genera such as ''Erythrosuchus''. In ''Fugusuchus'', the tooth row of the upper jaw extends beneath the orbit, or eye socket. This feature distinguishes it from more advanced erythrosuchids such as ''Garjainia ''Garjainia'' is an extinct genus of erythrosuchid archosauriform reptile from the Olenekian of Russia and South Africa. It was approximately 1.50–2 m (5–6 ft 8 in) long. It contained two species, ''Garjainia ...
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Middle Triassic
In the geologic timescale, the Middle Triassic is the second of three epochs of the Triassic period or the middle of three series in which the Triassic system is divided in chronostratigraphy. The Middle Triassic spans the time between Ma and Ma (million years ago). It is preceded by the Early Triassic Epoch and followed by the Late Triassic Epoch. The Middle Triassic is divided into the Anisian and Ladinian ages or stages. Formerly the middle series in the Triassic was also known as Muschelkalk. This name is now only used for a specific unit of rock strata with approximately Middle Triassic age, found in western Europe. Middle Triassic fauna Following the Permian–Triassic extinction event, the most devastating of all mass-extinctions, life recovered slowly. In the Middle Triassic, many groups of organisms reached higher diversity again, such as the marine reptiles (e.g. ichthyosaurs, sauropterygians, thallatosaurs), ray-finned fish and many invertebrate groups like ...
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Phylogenetic
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, protein amino acid sequences, or morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An unrooted tree diagram (a network) makes no assumption about the ancestral line, and does ...
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Gamosaurus
''Chasmatosuchus'' was an archosauriform reptile from the early Triassic period of European Russia. One of the earliest described archosauriforms, it was over 2 m long and is thought to have behaved like a modern crocodile. Its mouth had two distinct features: the top of its jaw hooked downwards to aid in holding prey, and the upper palate was lined with a row of teeth—a primitive feature lost in later archosaurs. ''Chasmatosuchus'' was formerly assigned to Proterosuchidae, but a 2016 cladistic analysis by Martin Ezcurra couldn't confidently place the species of ''Chasmatosuchus'' within Proterosuchidae. Instead their position remains unresolved due to the fragmentary nature of the known material, however ''Chasmatosuchus'' is more likely to be intermediate between proterosuchids and erythrosuchids and possibly closely related to '' Sarmatosuchus otschevi'' and '' Cuyosuchus huenei''. Ezcurra (2016) could only placed ''C. rossicus'' and ''C. magnus'' within ''Chasmatosuch ...
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Proterosuchia
Proterosuchia is one of the suborders of the paraphyletic group Thecodontia; containing the most primitive and ancestral forms. These were primitive, vaguely crocodile-like, archosauriforms that mostly lived during the Early Triassic epoch. The name Proterosuchia was coined by Robert Broom in 1906. In later classifications, several families were included, such as Proterosuchidae, Erythrosuchidae, and Proterochampsidae. Under the cladistic system, this is a paraphyletic grade, rather than a natural group. The name is therefore no longer used, although it can be found in many textbooks (up to and including Carroll's ''Vertebrate Paleontology and Evolution'') In current phylogenetic understanding, the Proterosuchia constitute the basal Archosauriformes Archosauriformes (Greek for 'ruling lizards', and Latin for 'form') is a clade of diapsid reptiles that developed from archosauromorph ancestors some time in the Latest Permian (roughly 252 million years ago). It was defined ...
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Erythrosuchia
Erythrosuchidae (meaning "red crocodiles" in Greek) are a family of large basal archosauriform carnivores that lived from the later Early Triassic (Olenekian) to the early Middle Triassic (Anisian). Naming The family Erythrosuchidae was named by David Meredith Seares Watson in 1917.D.M.S. Watson. 1917. "A sketch classification of the Pre-Jurassic tetrapod vertebrates". ''Proceedings of the Zoological Society of London'' 1917: 167–186 Description They were the apex predators of their day, with lengths of to almost . Their fossil remains are known to date from South Africa (Beaufort Group of the Karoo Basin), China, India and European Russia, from the Early to Middle Triassic. Erythrosuchids were unusually large and robust archosauromorphs. Several features set them apart from other archosauriformes and are also seen in later, more derived archosaurs. For example, they lack teeth on the palate, which are found in other early archosauriformes, such as ''Doswellia'' and euparker ...
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