Unenlagiinae
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Unenlagiinae
Unenlagiinae is a subfamily of long-snouted Paraves, paravian theropods. They are traditionally considered to be members of Dromaeosauridae, though some authors place them into their own family, Unenlagiidae, alongside the subfamily Halszkaraptorinae. Unenlagiines are known from South America, with possible unenlagiines being known from North America, Madagascar, Europe, and even Australia. Description Most unenlagiines have been discovered in Argentina. The largest was ''Austroraptor'', which measured up to 5–6 m (16.4–19.7 ft) in length, making it also one of the largest dromaeosaurids. The subfamily is distinguished from other dromaeosaurids by a tail stiffened by lengthy chevrons and superior processes, a reduced second pedal ungual, a posteriorly oriented pubis and very elongated snouts. Unenlagiines also had elongated, slender hindlimbs with a subarctometatarsalian metatarsus, which is characterized by the pinched metatarsal III at the upper end. Their distinct ...
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Dromaeosauridae
Dromaeosauridae () is a family of feathered theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek ('), meaning 'runner', and ('), meaning 'lizard'. In informal usage, they are often called raptors (after ''Velociraptor''), a term popularized by the film '' Jurassic Park''; a few types include the term "raptor" directly in their name and have come to emphasize their bird-like appearance and speculated bird-like behavior. Dromaeosaurid fossils have been found across the globe in North America, Europe, Africa, Asia, South America and Antarctica, with some fossils giving credence to the possibility that they inhabited Australia as well. They first appeared in the mid-Jurassic Period (late Bathonian stage, about 167 million years ago) and survived until the end of the Cretaceous (Maastrichtian stage, 66 ma), existing until the Cretaceous–Paleogene ex ...
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Austroraptor
''Austroraptor'' ( ) is a genus of large dromaeosaurid dinosaur that lived during the Late Cretaceous Period in what is now Argentina. ''Austroraptor'' was a large-sized, moderately-built, ground-dwelling, bipedal carnivore, estimated at in length. It is one of the largest dromaeosaurids known, with only ''Achillobator'', ''Dakotaraptor'', and ''Utahraptor'' approaching or surpassing it in length. Discovery and naming The type specimen of ''Austroraptor cabazai'', holotype MML-195, was recovered in the Bajo de Santa Rosa locality of the Allen Formation, in Río Negro, Argentina. The specimen was collected in 2002 by the team of Fernando Emilio Novas of the '' Museo Argentino de Ciencias Naturales''. It consists of a fragmentary skeleton including parts of the skull, lower jaw, a few neck and torso vertebrae, some ribs, a humerus, and assorted bones from both legs. The specimen was prepared by Marcelo Pablo Isasi and Santiago Reuil. In 2008, the type species ''Austroraptor cab ...
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Austroraptor Restoration
''Austroraptor'' ( ) is a genus of large dromaeosaurid dinosaur that lived during the Late Cretaceous Period in what is now Argentina. ''Austroraptor'' was a large-sized, moderately-built, ground-dwelling, bipedal carnivore, estimated at in length. It is one of the largest dromaeosaurids known, with only ''Achillobator'', ''Dakotaraptor'', and ''Utahraptor'' approaching or surpassing it in length. Discovery and naming The type specimen of ''Austroraptor cabazai'', holotype MML-195, was recovered in the Bajo de Santa Rosa locality of the Allen Formation, in Río Negro (Argentina), Río Negro, Argentina. The specimen was collected in 2002 by the team of Fernando Emilio Novas of the ''Museo Argentino de Ciencias Naturales''. It consists of a fragmentary skeleton including parts of the skull, lower jaw, a few neck and torso vertebrae, some ribs, a humerus, and assorted bones from both legs. The specimen was prepared by Marcelo Pablo Isasi and Santiago Reuil. In 2008, the type spec ...
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Ypupiara
''Ypupiara'' (meaning "the one who lives in the water") is an extinct genus of unenlagiine theropod from the Late Cretaceous Serra da Galga Formation of Brazil.Brum, Arthur Souza, Pêgas, Rodrigo Vargas, Bandeira, Kamila Luisa Nogueira, Souza, Lucy Gomes de, Campos, Diogenes de Almeida, & Kellner, Alexander Wilhelm Armin. (2021). A new Unenlagiinae (Theropoda: Dromaeosauridae) from the Late Cretaceous of Brazil. https://doi.org/10.1002/spp2.1375 It was the first member of the Dromaeosauridae to be discovered in South America and the first member of the Unenlagiinae to be discovered, but not the first to be identified as such. The type and only species, ''Y. lopai'', is known solely from a specimen that was destroyed in a fire in 2018. Discovery and naming The holotype, DGM 921-R, a right maxilla and dentary (which was associated with a fish jaw), was discovered in a layer of the Late Cretaceous Serra da Galga Formation of Brazil. It was found by Alberto Lopa sometime in the 1950s ...
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Unenlagiidae
Unenlagiidae is a proposed family of eumaniraptoran paravians that includes the subfamilies Unenlagiinae and possibly Halszkaraptorinae. Fossils of both subfamilies have been found in both Gondwanan and Laurasian deposits. The biology of the group suggests that some members were semiaquatic specialists. Classification The family Unenlagiidae traditionally includes the same members as the previously named subfamily of Dromaeosauridae, Unenlagiinae, so Unenlagiidae was often seen as a synonym of Dromaeosauridae. However, since the 2010s, there have been subsequent studies that have questioned this placement, necessitating the revival of the family name. Some have placed unenlagiids as outside Dromaeosauridae, being the sister taxon or closely related to Avialae, while others have placed the newly recognized halszkaraptorines in the family, as basal deinonychosaurs outside Dromaeosauridae and Troodontidae Troodontidae is a clade of bird-like theropod dinosaurs. During most o ...
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Unenlagia Paynemili
''Unenlagia'' (meaning "half-bird" in Latinized Mapudungun) is a genus of dromaeosaurid theropod dinosaur that lived in South America during the Late Cretaceous period. The genus ''Unenlagia'' has been assigned two species: ''U. comahuensis'', the type species described by Novas and Puerta in 1997, and ''U. paynemili'', described by Calvo ''et al.'' in 2004. Discovery and naming In 1996 in the Neuquén province of Argentina a skeleton of a theropod was discovered in the Sierra del Portezuelo and reported the same year. In 1997 Fernando Emilio Novas and Pablo Puerta named and described ''Unenlagia comahuensis''. The generic name is derived from Mapuche ''uñùm'', 'bird', and ''llag'', 'half', in reference to the fact that the describers considered the species to be a link between birds and more basal theropods. The specific name refers to the Comahue, the region the find was made. The holotype specimen, MCF PVPH 78, was uncovered in layers of the Portezuelo Formation dating ...
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Unenlagia Comahuensis
''Unenlagia'' (meaning "half-bird" in Latinized Mapudungun) is a genus of dromaeosaurid theropod dinosaur that lived in South America during the Late Cretaceous period. The genus ''Unenlagia'' has been assigned two species: ''U. comahuensis'', the Type (biology), type species described by Novas and Puerta in 1997, and ''U. paynemili'', described by Calvo ''et al.'' in 2004. Discovery and naming In 1996 in the Neuquén province of Argentina a skeleton of a theropod was discovered in the Sierra del Portezuelo and reported the same year. In 1997 Fernando Emilio Novas and Pablo Puerta named and described ''Unenlagia comahuensis''. The generic name is derived from Mapuche ''uñùm'', 'bird', and ''llag'', 'half', in reference to the fact that the describers considered the species to be a link between birds and more basal theropods. The specific name (zoology), specific name refers to the Comahue, the region the find was made. The holotype specimen, MCF PVPH 78, was uncovered in lay ...
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Pyroraptor
''Pyroraptor'' (meaning "fire thief") is an extinct genus of paravian dinosaur, probably a dromaeosaurid or unenlagiid (considering that unenlagiids are a separate family), from the Late Cretaceous Ibero-Armorican island, of what is now southern France and northern Spain. It lived during the late Campanian and early Maastrichtian stages, approximately 70.6 million years ago. It is known from a single partial specimen that was found in Provence in 1992, after a forest fire. The animal was named ''Pyroraptor olympius'' by Allain and Taquet in 2000. Discovery and naming Naming and Material The first remains of ''Pyroraptor olympius'', or P. olympius, were discovered in southeastern France, at the La Boucharde locality of the Arc Basin in Provence. It was described and named by French paleontologists Ronan Allain and Philippe Taquet in 2000, the type species and so far the only species is ''Pyroraptor olympius''. The genus name is Greek for "Fire thief", due to its remains ...
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Rahonavis
''Rahonavis'' is a genus of bird-like theropods from the Late Cretaceous (Maastrichtian, about 70 mya) of what is now northwestern Madagascar. It is known from a partial skeleton ( UA 8656) found by Catherine Forster and colleagues in Maevarano Formation rocks at a quarry near Berivotra, Mahajanga Province.Tudge, Colin (2009) ''The Bird:A Natural History of Who Birds Are, Where They Came From, and How They Live'/ref> ''Rahonavis'' was a small predator, at about long and 0.45-2.27 kg (1-5 lbs),Holtz, Thomas R. Jr. (2008) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages'Supplementary Information/ref> with the typical ''Velociraptor''-like raised sickle claw on the second toe. It was originally the first African coelurosaur until the Nqwebasaurus was discovered in 2000. The name ''Rahonavis'' means, approximately, "cloud menace bird", from Malagasy ' (RA-hoo-na, "cloud" or "menace") + Latin ' "bird". The specific name, ''R. ostromi'', was co ...
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Unenlagia
''Unenlagia'' (meaning "half-bird" in Latinized Mapudungun) is a genus of dromaeosaurid theropod dinosaur that lived in South America during the Late Cretaceous period. The genus ''Unenlagia'' has been assigned two species: ''U. comahuensis'', the Type (biology), type species described by Novas and Puerta in 1997, and ''U. paynemili'', described by Calvo ''et al.'' in 2004. Discovery and naming In 1996 in the Neuquén province of Argentina a skeleton of a theropod was discovered in the Sierra del Portezuelo and reported the same year. In 1997 Fernando Emilio Novas and Pablo Puerta named and described ''Unenlagia comahuensis''. The generic name is derived from Mapuche ''uñùm'', 'bird', and ''llag'', 'half', in reference to the fact that the describers considered the species to be a link between birds and more basal theropods. The specific name (zoology), specific name refers to the Comahue, the region the find was made. The holotype specimen, MCF PVPH 78, was uncovered in lay ...
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Halszkaraptor
''Halszkaraptor'' (; meaning " Halszka's seizer") is a genus of waterfowl-like dromaeosaurid dinosaurs from Mongolia that lived during the Late Cretaceous period. It contains only one known species, ''Halszkaraptor escuilliei''. The type specimen (holotype) has been compared to the bones of extant crocodilians and aquatic birds, and found evidence of a semiaquatic lifestyle, while some researchers question about semiaquatic ecology. A phylogenetic analysis revealed it was a member of the basal subfamily Halszkaraptorinae, along with ''Mahakala'' and ''Hulsanpes''. History of discovery The holotype specimen of ''Halszkaraptor'' likely came from the Djadochta Formation at Ukhaa Tolgod in southern Mongolia, and was illegally removed by fossil poachers. The fossil found its way to Japan and Great Britain, being owned by several collectors for some years until the Eldonia company of fossil dealer François Escuillié obtained it. He identified it as a new species, and in 2015 ...
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Eudromaeosauria
Eudromaeosauria ("true dromaeosaurs") is a subgroup of terrestrial dromaeosaurid theropod dinosaurs. They were relatively small to medium-sized, feathered hypercarnivores (with diets consisting almost entirely of other terrestrial vertebrates) that flourished in the Cretaceous Period. Eudromaeosaur fossils are known almost exclusively from the northern hemisphere. They first appeared in the early Cretaceous Period (early Aptian stage, about 124 million years ago) and survived until the end of the Cretaceous (Maastrichtian stage, Ma). The earliest known definitive eudromaeosaur is the dromaeosaurine '' Utahraptor ostrommaysi'', from the Cedar Mountain Formation, dated to 124 million years ago. However, the earlier (143-million-year-old) fossils such as those of '' Nuthetes destructor'' and several indeterminate teeth dating to the Kimmeridgian stage may represent eudromaeosaurs. Description While other dromaeosaurids filled a variety of specialized ecological niches, mainly tho ...
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