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Ustia (therapsid)
''Ustia'' is an extinct genus of biarmosuchian therapsids from the Middle Permian of Russia. It is known from a single species, ''Ustia atra'', which was described in 2003 from an isolated lower jaw. ''Ustia'' was classified in the family Ictidorhinidae, which also includes the genus ''Ictidorhinus'' from South Africa. Both are relatively small biarmosuchians. Several other Russian therapsids known only from lower jaw bones have been placed in Ictidorhinidae, and the family is likely a paraphyletic assemblage representing a small body type than a true clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, .... References Biarmosuchians Prehistoric therapsid genera Guadalupian synapsids of Europe Fossil taxa described in 2003 {{paleo-therapsid-stub ...
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Middle Permian
The Guadalupian is the second and middle series/epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson's Extinction and a significant mass extinction called the end-Capitanian extinction event. The Guadalupian was previously known as the Middle Permian. Name and background The Guadalupian is the second and middle series or epoch of the Permian. Previously called Middle Permian, the name of this epoch is part of a revision of Permian stratigraphy for standard global correlation. The name "Guadalupian" was first proposed in the early 1900s, and approved by the International Subcommission on Permian Stratigraphy in 1996. References to the Middle Permian still exist. The Guadalupian was preceded by the Cisuralian and followed by the Lop ...
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Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus comes above species and below family (taxonomy), family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants ...
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Biarmosuchia
Biarmosuchians are an extinct clade of non-mammalian synapsids from the Permian. They are the most basal group of the therapsids. All of them were moderately-sized, lightly-built carnivores, intermediate in form between basal sphenacodont "pelycosaurs" and more advanced therapsids. Biarmosuchians were rare components of Permian ecosystems, and the majority of species belong to the clade Burnetiamorpha, which are characterized by elaborate cranial ornamentation. Characteristics The biarmosuchian skull is very similar to the sphenacodontid skull, differing only in the larger temporal fenestra (although these are still small relative to later therapsids), slightly backward-sloping occiput (the reverse of the pelycosaur condition), reduced number of teeth, and single large canine teeth in both upper and lower jaws, and other features (Carroll 1988 pp. 370, Benton 2000 p. 114). In later specialised Biarmosuchia, these resemble the enlarged canines of the Gorgonopsia. The pr ...
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Therapsid
Therapsida is a major group of eupelycosaurian synapsids that includes mammals, their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more underneath the body, as opposed to the sprawling posture of many reptiles and salamanders. Therapsids evolved from "pelycosaurs", specifically within the Sphenacodontia, more than 279.5 million years ago. They replaced the "pelycosaurs" as the dominant large land animals in the Middle Permian through to the Early Triassic. In the aftermath of the Permian–Triassic extinction event, therapsids declined in relative importance to the rapidly diversifying reptiles during the Middle Triassic. The therapsids include the cynodonts, the group that gave rise to mammals ( Mammaliaformes) in the Late Triassic, around 225 million years ago. Of the non-mammalian therapsids, only cynodonts survived beyond the end of the Triassic, with the only other ...
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Ictidorhinidae
''Ictidorhinus'' is an extinct genus of biarmosuchian therapsids. Fossils have been found from the ''Dicynodon'' Assemblage Zone of the Beaufort Group in the Karoo Basin, South Africa and are of Late Permian age. It had a short snout and proportionally large orbits. These characteristics may be representative of a juvenile animal, possibly of ''Lycaenodon''. However, these two genera are not known to have existed at the same time, making it unlikely for ''Ictidorhinus'' material to be from a juvenile form of ''Lycaenodon''. Sidor, C. A. and Rubidge, B. S. (2006). ''Herpetoskylax hopsoni'', a new biarmosuchian (Therapsida: Biarmosuchia) from the Beaufort Group in South Africa. ''In:'' Matthew T. Carrano, Timothy J. Gaudin, Richard W. Blob and John R. Wible, eds., ''Amniote Paeobiology''. Chicago. University of Chicago Press. pp. 96. ''Ictidorhinus'' is the best-known representative of the family Ictidorhinidae, named by South African paleontologist Robert Broom in 1932. Many biarm ...
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Ictidorhinus
''Ictidorhinus'' is an extinct genus of biarmosuchian therapsids. Fossils have been found from the ''Dicynodon'' Assemblage Zone of the Beaufort Group in the Karoo Basin, South Africa and are of Late Permian age. It had a short snout and proportionally large orbits. These characteristics may be representative of a juvenile animal, possibly of ''Lycaenodon''. However, these two genera are not known to have existed at the same time, making it unlikely for ''Ictidorhinus'' material to be from a juvenile form of ''Lycaenodon''. Sidor, C. A. and Rubidge, B. S. (2006). ''Herpetoskylax hopsoni'', a new biarmosuchian (Therapsida: Biarmosuchia) from the Beaufort Group in South Africa. ''In:'' Matthew T. Carrano, Timothy J. Gaudin, Richard W. Blob and John R. Wible, eds., ''Amniote Paeobiology''. Chicago. University of Chicago Press. pp. 96. ''Ictidorhinus'' is the best-known representative of the family Ictidorhinidae, named by South African paleontologist Robert Broom in 1932. Many biarm ...
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Paraphyletic
In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic group (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of Synapomorphy and apomorphy, synapomorphies and symplesiomorphy, symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term was coined by Willi Hennig to apply to well-known taxa like Reptilia (reptiles) which, as commonly named and traditionally defined, is paraphyletic with respect to mammals and birds. Reptilia contains the last common ancestor of reptiles a ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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Biarmosuchians
Biarmosuchians are an extinct clade of non-mammalian synapsids from the Permian. They are the most basal group of the therapsids. All of them were moderately-sized, lightly-built carnivores, intermediate in form between basal sphenacodont "pelycosaurs" and more advanced therapsids. Biarmosuchians were rare components of Permian ecosystems, and the majority of species belong to the clade Burnetiamorpha, which are characterized by elaborate cranial ornamentation. Characteristics The biarmosuchian skull is very similar to the sphenacodontid skull, differing only in the larger temporal fenestra (although these are still small relative to later therapsids), slightly backward-sloping occiput (the reverse of the pelycosaur condition), reduced number of teeth, and single large canine teeth in both upper and lower jaws, and other features (Carroll 1988 pp. 370, Benton 2000 p. 114). In later specialised Biarmosuchia, these resemble the enlarged canines of the Gorgonopsia. The pre ...
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Prehistoric Therapsid Genera
Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of symbols, marks, and images appears very early among humans, but the earliest known writing systems appeared 5000 years ago. It took thousands of years for writing systems to be widely adopted, with writing spreading to almost all cultures by the 19th century. The end of prehistory therefore came at very different times in different places, and the term is less often used in discussing societies where prehistory ended relatively recently. In the early Bronze Age, Sumer in Mesopotamia, the Indus Valley Civilisation, and ancient Egypt were the first civilizations to develop their own scripts and to keep historical records, with their neighbors following. Most other civilizations reached the end of prehistory during the following Iron Age. T ...
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Guadalupian Synapsids Of Europe
The Guadalupian is the second and middle series/epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson's Extinction and a significant mass extinction called the end-Capitanian extinction event. The Guadalupian was previously known as the Middle Permian. Name and background The Guadalupian is the second and middle series or epoch of the Permian. Previously called Middle Permian, the name of this epoch is part of a revision of Permian stratigraphy for standard global correlation. The name "Guadalupian" was first proposed in the early 1900s, and approved by the International Subcommission on Permian Stratigraphy in 1996. References to the Middle Permian still exist. The Guadalupian was preceded by the Cisuralian and followed by the Lop ...
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