Triplagnostus
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Triplagnostus
''Ptychagnostus'' is a member of the agnostida that lived during the Cambrian period. Ptychagnostidae generally do not exceed one centimetre in length. Their remains are rarely found in empty tubes of the polychaete worm ''Selkirkia''. The genus probably ranged throughout the water column. It has two glabellar lobes, and three pygidial lobes. Type species ''Agnostus punctuosus'' Angelin, 1851 from the ''Pt. punctuosus'' Zone of the Alum Shale (Drumian), Sweden (by original designation). Official ruling on the conservation of accepted usage of ''A. punctuosus'' as the type species was given by the International Commission on Zoological Nomenclature, 1993. Remarks ''Ptychagnostus affinis'' (Brøgger 1878) was once considered a subspecies of ''Pt. punctuosus''. Laurie (2008) grouped ''punctuosus'' and ''affinis'' within ''Ptychagnostus'', but preferred to place the closely related ''atavus'' within ''Acidusus''. Ptychagnostidae Genera Høyberget & Bruton (2008) HØYBERGET M. & ...
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Otto Jaekel
Otto Max Johannes Jaekel (21 February 1863 – 6 March 1929) was a German paleontologist and geologist. Biography Jaekel was born in Neusalz (Nowa Sól), Prussian Silesia, the son of a builder and the youngest of seven children. He studied at the ''Ritterakademie'' in Liegnitz (Legnica). After graduating in 1883, he came to study geology and paleontology under Ferdinand Roemer in Breslau (Wrocław) until 1885. Karl von Zittel awarded a PhD to Jaekel in Munich in 1886. Between 1887 and 1889, Jaekel was an assistant of E.W. Benecke at the Geologisch-Paläontologisches Institut in Straßburg, where he received his Habilitation. He worked at the Friedrich-Wilhelms-Universität Berlin and at the Geologisch-Paläontologisches Museum (a combined post) from 1894. Jaekel was considered as an ordinary professor of geology at the University of Vienna in 1903, but this was blocked by intrigue. Between 1906 and 1928, Jaekel was a professor at the University of Greifswald, where he fo ...
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Agnostida
Agnostida is an order of arthropod which have classically been seen as a group of highly modified trilobites, though some recent research has doubted this placement. Regardless, they appear to be close relatives as part of the Artiopoda. They are present in the Lower Cambrian fossil record along with trilobites from the Redlichiida, Corynexochida, and Ptychopariida orders, and were highly diverse throughout the Cambrian. Agnostidan diversity severely declined during the Cambrian-Ordovician transition, and the last agnostidans went extinct in the Late Ordovician. Systematics The Agnostida are divided into two suborders — Agnostina and Eodiscina — which are then subdivided into a number of families. As a group, agnostids are isopygous, meaning their pygidium is similar in size and shape to their cephalon. Most agnostid species were eyeless. The systematic position of the order Agnostida within the class Trilobita remains uncertain, and there has been continuing deb ...
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Cambrian
The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Ca ...
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Selkirkia
''Selkirkia'' is a genus of predatory, tubicolous priapulid worms known from the Middle Cambrian Burgess Shale, Ogygopsis Shale and Puncoviscana Formation. 142 specimens of ''Selkirkia'' are known from the Greater Phyllopod bed, where they comprise 0.27% of the community. In the Burgess Shale, 20% of the tapering, organic-walled tubes are preserved with the worm inside them, whereas the other 80% are empty (or sometimes occupied by one or more small agnostid trilobites). Whilst alive, the tubes were probably vertical, whereas trilobite-occupied tubes are horizontal. Morphology ''Selkirkia'' had a body divisible into a proboscis towards the anterior of a trunk enclosed by a tube. The proboscis would have been partially invertable and was armed with several spinules and spines, decreasing size distally overall. It was controlled by at least two sets of anterior retractor muscles. Immediately behind the proboscis was the trunk, smooth for the most part but lined with papillae tow ...
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Richard A
Richard is a male given name. It originates, via Old French, from Old Frankish and is a compound of the words descending from Proto-Germanic ''*rīk-'' 'ruler, leader, king' and ''*hardu-'' 'strong, brave, hardy', and it therefore means 'strong in rule'. Nicknames include "Richie", "Dick", "Dickon", " Dickie", "Rich", "Rick", "Rico", "Ricky", and more. Richard is a common English, German and French male name. It's also used in many more languages, particularly Germanic, such as Norwegian, Danish, Swedish, Icelandic, and Dutch, as well as other languages including Irish, Scottish, Welsh and Finnish. Richard is cognate with variants of the name in other European languages, such as the Swedish "Rickard", the Catalan "Ricard" and the Italian "Riccardo", among others (see comprehensive variant list below). People named Richard Multiple people with the same name * Richard Andersen (other) * Richard Anderson (other) * Richard Cartwright (other) * Ri ...
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Drumian
The Drumian is a stage of the Miaolingian Series of the Cambrian. It succeeds the Wuliuan and precedes the Guzhangian. The base is defined as the first appearance of the trilobite ''Ptychagnostus atavus'' around million years ago. The top is defined as the first appearance of another trilobite ''Lejopyge laevigata'' around million years ago. The GSSP is defined in the ''Drumian section'' () in the Drum Mountains, Millard County, Utah, United States. The stage was also named after the Drum Mountains. The section is an outcrop of the Wheeler Formation The Wheeler Shale (named by Charles Walcott) is a Cambrian ( 507  Ma) fossil locality world-famous for prolific agnostid and ''Elrathia kingii'' trilobite remains (even though many areas are barren of fossils) and represents a Konzent ..., a succession of calcareous shales. The precise base of the Drumian is a laminated limestone above the base of the Wheeler Formation. References Cambrian geochronology Geo ...
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Ptychagnostus Atavus
''Ptychagnostus atavus'' is a species of agnostid trilobite. It was originally described by Swedish paleontologist Sven Axel Tullberg as ''Agnostus atavus'' in 1880. It is used in biostratigraphy as an index fossil. Its first appearance at the GSSP section in the Wheeler Shale of Utah is defined as the beginning of the Drumian Age (around million years ago) of the Miaolingian (Middle Cambrian Middle or The Middle may refer to: * Centre (geometry), the point equally distant from the outer limits. Places * Middle (sheading), a subdivision of the Isle of Man * Middle Bay (other) * Middle Brook (other) * Middle Creek (di ...). Laurie (2008) LAURIE J. R. 2008. Species relationships in the Ptychagnostidae (Cambrian, Agnostina). In: Ra´bano I., Gozalo R. & GarciaBellido D. eds. ''Advances in trilobite research'', pp. 211–218. Cuadernos del Museo Geominero 9. grouped ''punctuosus'' and ''affinis'' within ''Ptychagnostus'', but preferred to place the closely re ...
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Agnostida Genera
Agnostida is an order of arthropod which have classically been seen as a group of highly modified trilobites, though some recent research has doubted this placement. Regardless, they appear to be close relatives as part of the Artiopoda. They are present in the Lower Cambrian fossil record along with trilobites from the Redlichiida, Corynexochida, and Ptychopariida orders, and were highly diverse throughout the Cambrian. Agnostidan diversity severely declined during the Cambrian-Ordovician transition, and the last agnostidans went extinct in the Late Ordovician. Systematics The Agnostida are divided into two suborders — Agnostina and Eodiscina — which are then subdivided into a number of families. As a group, agnostids are isopygous, meaning their pygidium is similar in size and shape to their cephalon. Most agnostid species were eyeless. The systematic position of the order Agnostida within the class Trilobita remains uncertain, and there has been continuing ...
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Cambrian Trilobites
The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Cambrian bi ...
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Burgess Shale Fossils
The fossils of the Burgess Shale, like the Burgess Shale itself, formed around 505 million years ago in the Mid Cambrian period. They were discovered in Canada in 1886, and Charles Doolittle Walcott collected over 65,000 specimens in a series of field trips up to the alpine site from 1909 to 1924. After a period of neglect from the 1930s to the early 1960s, new excavations and re-examinations of Walcott's collection continue to reveal new species, and statistical analysis suggests that additional discoveries will continue for the foreseeable future. Stephen Jay Gould's book '' Wonderful Life'' describes the history of discovery up to the early 1980s, although his analysis of the implications for evolution has been contested. The fossil beds are in a series of shale layers, averaging and totalling about in thickness. These layers were deposited against the face of a high undersea limestone cliff. All these features were later raised up above current sea level during the cr ...
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Wheeler Shale
The Wheeler Shale (named by Charles Walcott) is a Cambrian ( 507  Ma) fossil locality world-famous for prolific agnostid and ''Elrathia kingii'' trilobite remains (even though many areas are barren of fossils) and represents a Konzentrat-Lagerstätte. Varied soft bodied organisms are locally preserved, a fauna (including ''Naraoia'', ''Wiwaxia'' and ''Hallucigenia'') and preservation style (carbonaceous film) normally associated with the more famous Burgess Shale. As such, the Wheeler Shale also represents a Konservat-Lagerstätten. Together with the Marjum Formation and lower Weeks Formation, the Wheeler Shale forms of limestone and shale exposed in one of the thickest, most fossiliferous and best exposed sequences of Middle Cambrian rocks in North America. At the type locality of Wheeler Amphitheater, House Range, Millard County, western Utah, the Wheeler Shale consists of a heterogeneous succession of highly calcareous shale, shaley limestone, mudstone and thi ...
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