''Ptychagnostus atavus'' is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

agnostid Agnostidae is a family of Agnostida trilobites. Like all Agnostina, they were eyeless and bore only two thoracic segments. They ranged in benthic waters across the globe from 508 to 461 million years ago, containing the following genera, among o ...

trilobite Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the At ...

. It was originally described by

Swedish Swedish or ' may refer to: Anything from or related to Sweden, a country in Northern Europe. Or, specifically: * Swedish language, a North Germanic language spoken primarily in Sweden and Finland ** Swedish alphabet, the official alphabet used by ...

paleontologist Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...

Sven Axel Tullberg as ''Agnostus atavus'' in 1880. It is used in

biostratigraphy Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock Stratum, strata by using the fossil assemblages contained within them.Hine, Robert. “Biostratigraphy.” ''Oxford Reference: Dictiona ...

as an

index fossil Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock Stratum, strata by using the fossil assemblages contained within them.Hine, Robert. “Biostratigraphy.” ''Oxford Reference: Dictiona ...

. Its

first appearance In American comic books and other stories with a long history, first appearance refers to the first issue to feature a fictional character. These issues are often highly valued by collectors due to their rarity and iconic status. Reader intere ...

at the

GSSP A Global Boundary Stratotype Section and Point (GSSP) is an internationally agreed upon reference point on a stratigraphic section which defines the lower boundary of a stage on the geologic time scale. The effort to define GSSPs is conducted b ...

section in the

Wheeler Shale The Wheeler Shale (named by Charles Walcott) is a Cambrian ( 507 Ma) fossil locality world-famous for prolific agnostid and ''Elrathia kingii'' trilobite remains (even though many areas are barren of fossils) and represents a Konzent ...

Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...

is defined as the beginning of the

Drumian The Drumian is a stage of the Miaolingian Series of the Cambrian. It succeeds the Wuliuan and precedes the Guzhangian. The base is defined as the first appearance of the trilobite ''Ptychagnostus atavus'' around million years ago. The top is def ...

Age Age or AGE may refer to: Time and its effects * Age, the amount of time someone or something has been alive or has existed ** East Asian age reckoning, an Asian system of marking age starting at 1 * Ageing or aging, the process of becoming older ...

(around

million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...

) of the

Miaolingian The Miaolingian is the third Series of the Cambrian Period, and was formally named in 2018. It lasted from about to million years ago and is divided in ascending order into 3 stages: the Wuliuan, Drumian, and Guzhangian. The Miaolingian is prec ...

(

Middle Cambrian Middle or The Middle may refer to: * Centre (geometry), the point equally distant from the outer limits. Places * Middle (sheading), a subdivision of the Isle of Man * Middle Bay (disambiguation) * Middle Brook (disambiguation) * Middle Creek (di ...

). Laurie (2008) LAURIE J. R. 2008. Species relationships in the Ptychagnostidae (Cambrian, Agnostina). In: Ra´bano I., Gozalo R. & GarciaBellido D. eds. ''Advances in trilobite research'', pp. 211–218. Cuadernos del Museo Geominero 9. grouped ''punctuosus'' and ''affinis'' within ''Ptychagnostus'', but preferred to place the closely related ''atavus'' within ''Acidusus''.

References

Agnostoidea Cambrian trilobites Fossil taxa described in 1880 Index fossils Drumian Wheeler Shale Drumian species {{agnostida-stub