Trigonotarbid
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Trigonotarbid
The order Trigonotarbida is a group of extinct arachnids whose fossil record extends from the late Silurian to the early Permian (Pridoli epoch, Pridoli to Sakmarian).Dunlop, J. A., Penney, D. & Jekel, D. 2020A summary list of fossil spiders and their relatives In World Spider Catalog. Natural History Museum Bern, online at http://wsc.nmbe.ch, version 20.5 These animals are known from several localities in Europe and North America, as well as a single record from Argentina. Trigonotarbids can be envisaged as spider-like arachnids, but without silk-producing spinnerets. They ranged in size from a few millimetres to a few centimetres in body length and had segmented abdomens (opisthosoma), with the dorsal exoskeleton (Tergum, tergites) across the backs of the animals' abdomens, which were characteristically divided into three or five separate plates. Probably living as predators on other arthropods, some later trigonotarbid species were quite heavily armoured and protected themselves ...
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Arthropod
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arthropod cuticle, cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an exoskeleton, external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior Organ (anatomy), organs; it has an open circulatory system. Like their exteriors, the internal or ...
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Maiocercus
''Maiocercus celticus'' is a species of early Trigonotarbida, trigonotarbid arachnid from the Upper Carboniferous of Westhoughton, Lancashire, UK. The species was first described in 1902, with a "new species" being described in 1911 (''M. orbicularis'') which has been proven as being a junior synonym of ''M. celticus''. ''M. celticus'' is the type species of the genus ''Maiocercus''. Originally zoologist Reginald Innes Pocock compared ''M. celticus'' to Brachypyge, with later evidence showing that ''Brachypyge'' had "opisthosoma which were much longer than wide; with the pleural laminæ of the second and third pleura-bearing terga being inclined slightly backwards" (''Brachypyge'') with ''Maiocercus'' having the “opisthosoma much wider than long; the pleural laminæ of the first, second, third, and fourth sterna being inclined slightly forwards”. The original drawing which showed ''Maiocercus'' described a pitting on the underside of the slightly forwarded laminæ, with a ...
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Spider
Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species diversity among all orders of organisms. Spiders are found worldwide on every continent except for Antarctica, and have become established in nearly every land habitat. , 50,356 spider species in 132 families have been recorded by taxonomists. However, there has been debate among scientists about how families should be classified, with over 20 different classifications proposed since 1900. Anatomically, spiders (as with all arachnids) differ from other arthropods in that the usual body segments are fused into two tagmata, the cephalothorax or prosoma, and the opisthosoma, or abdomen, and joined by a small, cylindrical pedicel, however, as there is currently neither paleontological nor embryological evidence that spiders ever had ...
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Palaeocharinus
''Palaeocharinus'' is a genus of extinct trigonotarbid arachnids known from the Devonian of western Europe. The genus was first found and described in the Rhynie chert in the 1920s by Arthur Stanley Hirst and S. Maulik. The family to which the genus belongs may be paraphyletic. Species * ''Palaeocharinus calmani'' (Hirst, 1923) – Early Devonian The Early Devonian is the first of three Epoch (geology), epochs comprising the Devonian period, corresponding to the Lower Devonian Series (stratigraphy), series. It lasted from and began with the Lochkovian Stage , which was followed by the P ..., Scotland * ''Palaeocharinus hornei'' (Hirst, 1923) – Early Devonian, Scotland * ''Palaeocharinus kidstoni'' (Hirst, 1923) – Early Devonian, Scotland * ''Palaeocharinus rhyniensis'' (Hirst, 1923) – Early Devonian, Scotland * ''Palaeocharinus scourfieldi'' (Hirst, 1923) – Early Devonian, Scotland * ''Palaeocharinus tuberculatus'' (Fayers, Dunlop & Trewin, 2005) – Early Devonian ...
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Late Silurian
The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods (myriapods, arachnids and hexapods) became fully terrestrialized. A significant evolutionary milestone during t ...
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Eophrynidae
Eophrynidae is a family of the extinct arachnid order Trigonotarbida The order Trigonotarbida is a group of extinct arachnids whose fossil record extends from the late Silurian to the early Permian ( Pridoli to Sakmarian).Dunlop, J. A., Penney, D. & Jekel, D. 2020A summary list of fossil spiders and their relative .... Eophrynids lived during the Carboniferous period in what is now modern Europe and North America. The family is probably found within the "eophrynid assemblage" clade: (''Aphantomartus'' (''Alkenia'' (''Pseudokreischeria'' (''Kreischeria'' (''Eophrynus'' + ''Pleophrynus''))))). Genera *'' Areomartus'' Petrunkevitch, 1913 *'' Eophrynus'' Woodward, 1871 *'' Nyranytarbus'' Harvey & Selden, 1995 *'' Petrovicia'' Frič, 1904 *'' Planomartus'' Petrunkevitch, 1953 *'' Pleophrynus'' Petrunkevitch, 1945 *'' Pocononia'' Petrunkevitch, 1953 *'' Somaspidion'' Jux, 1982 *'' Stenotrogulus'' Frič, 1904 *'' Vratislavia'' Frič, 1904 References Trigonotarbid ...
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Anthracomartidae
Anthracomartidae, first described by Haase, 1890, is a family of the extinct arachnid order Trigonotarbida The order Trigonotarbida is a group of extinct arachnids whose fossil record extends from the late Silurian to the early Permian ( Pridoli to Sakmarian).Dunlop, J. A., Penney, D. & Jekel, D. 2020A summary list of fossil spiders and their relative .... It is likely to be most closely related to the Archaeomartidae, based on a 2014 cladistic analysis, within the clade (Palaeocharinus (Archaeomartidae + Anthracomartidae)). Genera *'' Anthracomartus'' Karsch, 1882 *'' Brachylycosa'' Frič, 1904 *'' Brachypyge'' Woodward, 1878 *'' Cleptomartus'' Petrunkevitch, 1949 *'' Coryphomartus'' Petrunkevitch, 1945 *'' Cryptomartus'' Petrunkevitch, 1945 *'' Maiocercus'' Pocock, 1911 *'' Oomartus'' Petrunkevitch, 1953 *'' Pleomartus'' Petrunkevitch, 1945 *'' Promygale'' Frič, 1904 References Trigonotarbids Paleozoic arachnids Silurian arachnids Devonian arachnids ...
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Silurian
The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods (myriapods, arachnids and hexapods) became fully terrestrialized. A significant evolutionary milestone during ...
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Europe
Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia and it is located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. Comprising the westernmost peninsulas of Eurasia, it shares the continental landmass of Afro-Eurasia with both Africa and Asia. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south and Asia to the east. Europe is commonly considered to be Boundaries between the continents of Earth#Asia and Europe, separated from Asia by the drainage divide, watershed of the Ural Mountains, the Ural (river), Ural River, the Caspian Sea, the Greater Caucasus, the Black Sea and the waterways of the Turkish Straits. "Europe" (pp. 68–69); "Asia" (pp. 90–91): "A commonly accepted division between Asia and E ...
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Sakmarian
In the geologic timescale, the Sakmarian is an age or stage of the Permian. It is a subdivision of the Cisuralian Epoch or Series. The Sakmarian lasted between 293.52 and million years ago (Ma). It was preceded by the Asselian and followed by the Artinskian.; 2004: ''A Geologic Time Scale 2004'', Cambridge University Press Stratigraphy The Sakmarian Stage is named after the Sakmara River in the Ural Mountains, a tributary to the Ural River. The stage was introduced into scientific literature by Alexander Karpinsky in 1874. In Russian stratigraphy, it originally formed a substage of the Artinskian Stage. Currently, the ICS uses it as an independent stage in its international geologic timescale. The base of the Sakmarian Stage is laid with the first appearance of conodont species ''Streptognathodus postfusus'' in the fossil record. A global reference profile for the base (a GSSP), located in the southern Ural Mountains, Russia, was ratified in 2018. The top of the Sakmarian (the ...
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Spinneret
A spinneret is a silk-spinning organ of a spider or the larva of an insect. Some adult insects also have spinnerets, such as those borne on the forelegs of Embioptera. Spinnerets are usually on the underside of a spider's opisthosoma, and are typically segmented. While most spiders have six spinnerets, some have two, four, or eight. They can move both independently and in concert. Most spinnerets are not simple structures with a single orifice producing a single thread, but complex structures of many microscopic spigots, each producing one filament. This produces the necessary orientation of the protein molecules, without which the silk would be weak and useless. Spigots can be singular or found in groups, which also permits spiders to combine multiple filaments in different ways to produce many kinds of silk for various purposes. Spinneret morphology can help arachnologists identify the taxon of a specimen and the specific morphology of a spigot can determine its use as well ...
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