Trematopidae
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Trematopidae
Trematopidae is a family of dissorophoid temnospondyl spanning the late Carboniferous to the early Permian. Together with Dissorophidae, the family forms Olsoniformes, a clade comprising the medium-large terrestrial dissorophoids. Trematopids are known from numerous localities in North America, primarily in New Mexico, Oklahoma, and Texas, and from the Bromacker quarry in Germany. History of study The clade Trematopidae was first proposed by American paleontologist S.W. Williston in 1910, although it was named as "Trematopsidae" following the historical (but inaccurate) derivation from the genus "''Trematops''" (now synonymized with ''Acheloma''). British paleontologist D.M.S. Watson proposed a related clade in 1919, Achelomidae, for ''Acheloma'', based on perceived differences separating the taxa; this is now considered a junior synonym of Trematopidae following guidelines of historical precedent. 19th century history In 1882, American paleontologist Edward Drinker Cope na ...
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Dissorophoidea
Dissorophoideans are a clade of medium-sized, temnospondyl amphibians that appeared during the Moscovian in Euramerica, and continued through to the Late Permian and the Early Triassic of Gondwana. They are distinguished by various details of the skull, and many forms seem to have been well adapted for life on land. Since 2008, Lissamphibia has been progressively widely considered part of this clade, but this position is still disputed by some authors. It is possible that the small Permo-Carboniferous Micromelerpetontidae and the large Late Permian Melosauridae may also belong in this clade. Phylogeny An extensive phylogenetic analysis of dissorophoids conducted in 2016 and 2018 found that the families Dissorophidae and Trematopidae are more closely related to each other than either is to the family Amphibamidae. Following a 2008 study, the Dissorophidae-Trematopidae clade was called Olsoniformes. Below is the cladogram from the 2018 analysis: References * Huttenlocker, A ...
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Tambachia
''Tambachia'' is an extinct genus of dissorophoid temnospondyl within the family Trematopidae. It is known from the Early Permian Tambach Formation (the lowermost unit of the Upper Rotliegend) near the town of Tambach-Dietharz in Thuringia, Germany. ''Tambachia'' is the first trematopid to have been discovered outside the United States. Discovery The holotype specimen of ''Tambachia trogallas'', known as MNG 7722, has been found from an outcrop of the Tambach Formation at the Bromacker locality in the Thuringian Forest of central Germany. It consists of a skull and much of the postcranial skeleton. The only major portion of the skeleton that is missing is the presacral vertebral column. The Bromacker locality is a sandstone quarry that is well known for tetrapod trackways and articulated skeletons of terrestrial and semiterrestrial amphibians and reptiles. MNG 7722 was found in red-bed fluvial deposits consisting of well consolidated mudstones in flat-bedded channel fills ...
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Temnospondyli
Temnospondyli (from Greek τέμνειν, ''temnein'' 'to cut' and σπόνδυλος, ''spondylos'' 'vertebra') is a diverse order of small to giant tetrapods—often considered primitive amphibians—that flourished worldwide during the Carboniferous, Permian, and Triassic periods. A few species continued into the Jurassic and Cretaceous periods. Fossils have been found on every continent. During about 210 million years of evolutionary history, they adapted to a wide range of habitats, including freshwater, terrestrial, and even coastal marine environments. Their life history is well understood, with fossils known from the larval stage, metamorphosis, and maturity. Most temnospondyls were semiaquatic, although some were almost fully terrestrial, returning to the water only to breed. These temnospondyls were some of the first vertebrates fully adapted to life on land. Although temnospondyls are considered amphibians, many had characteristics, such as scales and armour-like bo ...
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Fedexia Striegeli
''Fedexia'' is an extinct genus of carnivorous temnospondyl within the family Trematopidae. It lived 300 million years ago during the late Carboniferous period. It is estimated to have been long, and likely resembled a salamander.Pitt student finds fossil of ancient amphibian
Pittsburgh Post-Gazette, March 15, 2010 ''Fedexia'' is known from a single skull found in , . It is named after the shipping service FedEx, which owned the land where the



Anconastes
''Anconastes'' is an extinct genus of dissorophoid temnospondyl within the family Trematopidae. It is known from two specimens from the Late Carboniferous Cutler Formation of north-central New Mexico in the southwestern United States. The genus name derives from two Greek roots, ''ankos'' ("mountain glen or valley") and ''nastes'' ("inhabitant"), which refers to the type locality of El Cobre Canyon where the specimens were found. The specific name is derived from the Latin word ''vesperus'' ("western"). The more complete specimen, the holotype, is a partial skull with articulated mandibles and a substantial amount of the postcranial skeleton. The less complete specimen, the paratype, consists only of the right margin of the skull with an articulated mandible. Anatomy When originally described, only three other trematopid taxa were recognized: '' Acheloma, Actiobates'', and "'' Trematops''" (now a synonym of ''Acheloma''); '' Ecolsonia'', which is now considered a trematopid by ...
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Fedexia
''Fedexia'' is an extinct genus of carnivorous temnospondyl within the family Trematopidae. It lived 300 million years ago during the late Carboniferous period. It is estimated to have been long, and likely resembled a salamander.Pitt student finds fossil of ancient amphibian
Pittsburgh Post-Gazette, March 15, 2010 ''Fedexia'' is known from a single skull found in , . It is named after the shipping service FedEx, which owned the land where the

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Phonerpeton
''Phonerpeton'' is an extinct genus of dissorophoid temnospondyl within the family Trematopidae that is known from the early Permian The Permian ( ) is a geologic period and System (stratigraphy), stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last ... of Texas. History of study ''Phonerpeton'' was first named by Dilkes (1990), with ''Phonerpeton pricei'' as the only species. The genus name comes from the Greek words '''phonos''' (murderous) and '''herpeton''' (creeper). The holotype of ''P. pricei'' was previously described by Olson (1941) as ''Acheloma pricei''. Dilkes also synonymized this species with ''Acheloma whitei'', which was also described by Olson (1941). Material referred to the taxon comes from the Archer City, Nocona, and Petrolia Formations and was collected between 1934 and 1952 by parties led by A.S. Romer, L.I. Price, and ...
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Olsoniformes
Olsoniformes is a clade of dissorophoid temnospondyls. It includes the families Dissorophidae and Trematopidae. Most members of the clade were highly adapted to a terrestrial lifestyle. The clade was named in 2008 and is defined as the least inclusive clade containing ''Dissorophus multicinctus'' (a dissorophid) and ''Acheloma cumminsi'' (a trematopid) but not ''Amphibamus grandiceps, Micromelerpeton credneri,'' and ''Apateon pedestris''. Olsoniforms share various features such as a stout and low ilium and a thin cultriform process. Trematopids are known from the Late Carboniferous and the Early Permian of Europe and across much of North America, while dissorophids are primarily found in Early Permian deposits in the central United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated te ...
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Dissorophidae
Dissorophidae is an extinct family of medium-sized, temnospondyl amphibians that flourished during the late Carboniferous and early Permian periods. The clade is known almost exclusively from North America. History of study Dissorophidae is a diverse clade that was named in 1902 by George A. Boulenger. Junior synonyms include Otocoelidae, Stegopidae, and Aspidosauridae. Early in the study of dissorophoids when the relationships of different taxa were not well-resolved and most taxa had not been described, Dissorophidae sometimes came to include taxa that are now not regarded as dissorophids and may have excluded earlier described taxa that are now regarded as dissorophids. Amphibamiforms were widely regarded as small-bodied dissorophids, and at one point, Dissorophidae was also suggested to also include Trematopidae. 19th century In 1895, American paleontologist Edward Drinker Cope named '' Dissorophus'' from the early Permian of Texas. This was the first dissorophid to be d ...
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Acheloma
''Acheloma'' (also known as ''Trematops milleri'') is an extinct genus of temnospondyl that lived during the Early Permian. The type species is ''A. cumminsi''. History of study ''Acheloma'' was named by Edward Drinker Cope in 1882 based on a partial skull with associated postcranial elements from the Arroyo Formation of Texas; the specimen is currently reposited at the American Museum of Natural History in New York. Subsequent discoveries of large trematopids from the Arroyo Formation were named as different species of ''Trematops'' (''T. milleri, T. willistoni''), but these have since been synonymized with ''Acheloma cumminsi''. ''Trematops stonei'' from the Washington Formation of Ohio and ''Trematops thomasi'' from Oklahoma have also been synonymized with ''A. cumminsi''. A second species of ''Acheloma'' was described by Polley & Reisz (2011) from the Richards Spur locality in Oklahoma. Anatomy Schoch & Milner (2014) provide nine characters in their diagnosis of ''A ...
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Ecolsonia Cutlerensis
''Ecolsonia'' is an extinct genus of trematopid temnospondyl. Its phylogenetic position within Olsoniformes has been historically debated, but it is presently considered to be a trematopid. History of study The holotype of ''Ecolsonia'' is a partial skull that was collected in 1963 from the VanderHoof quarry in the Early Permian Cutler Formation of New Mexico. The specimen was originally reposited at the now defunct University of California, Los Angeles Vertebrate Paleontology collections. The genus name is for the American paleontologist Everett C. Olson, and the species name is for the geologic provenance. A number of more complete specimens from the Morfin quarry in the Cutler Formation were discovered in 1983 and described by Berman et al. (1985), including a complete skull and large amounts of postcranial material. These specimens are currently reposited at the Carnegie Museum of Natural History. Anatomy Schoch & Milner (2014) listed seven features in the diagnosis of ...
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Acheloma Cumminsi
''Acheloma'' (also known as ''Trematops milleri'') is an extinct genus of temnospondyl that lived during the Early Permian. The type species is ''A. cumminsi''. History of study ''Acheloma'' was named by Edward Drinker Cope in 1882 based on a partial skull with associated postcranial elements from the Arroyo Formation of Texas; the specimen is currently reposited at the American Museum of Natural History in New York. Subsequent discoveries of large trematopids from the Arroyo Formation were named as different species of ''Trematops'' (''T. milleri, T. willistoni''), but these have since been synonymized with ''Acheloma cumminsi''. ''Trematops stonei'' from the Washington Formation of Ohio and ''Trematops thomasi'' from Oklahoma have also been synonymized with ''A. cumminsi''. A second species of ''Acheloma'' was described by Polley & Reisz (2011) from the Richards Spur locality in Oklahoma. Anatomy Schoch & Milner (2014) provide nine characters in their diagnosis of ''A ...
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