Tholurna
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Tholurna
''Tholurna'' is a genus of lichenized fungi in the family Caliciaceae. The genus is monotypic, containing the single species ''Tholurna dissimilis''. This species was first described by Johannes Musaeus Norman as ''Podocratera dissimilis'' in 1861; he circumscribed In geometry, the circumscribed circle or circumcircle of a polygon is a circle that passes through all the vertices of the polygon. The center of this circle is called the circumcenter and its radius is called the circumradius. Not every polyg ... the new genus ''Tholurna'' the same year to contain the species. It is found in Scandinavia and western North America. References Caliciales Lichen genera Caliciales genera Taxa described in 1861 Monotypic Lecanoromycetes genera {{Caliciales-stub ...
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Caliciaceae
The Caliciaceae are a family of mostly lichen-forming fungi belonging to the class Lecanoromycetes in the division Ascomycota. Although the family has had its classification changed several times throughout its taxonomic history, the use of modern molecular phylogenetic methods have helped to establish its current placement in the order Caliciales. Caliciaceae contains 36 genera and about 600 species. The largest genus is ''Buellia'', with around 300 species; there are more than a dozen genera that contain only a single species. Most Caliciaceae grow on bark, dead wood, or rocks. Some members of this family, particularly those of the type genus, ''Calicium'', are characterized by the presence of thin-walled and short-lasting asci (spore-bearing cells) and a mazaedium, which is an accumulation of loose, maturing spores covering the surface of the fruiting body. The resulting passive spore dispersal is relatively rare amongst the Ascomycota. The mazaedium, usually black, either ...
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Caliciales
Caliciales is an order of mostly lichenized fungi in the class Lecanoromycetes. It consists of two families: Caliciaceae and Physciaceae, which together contain 54 genera and more than 1200 species. The order was circumscribed by American botanist Charles Edwin Bessey in 1907. Families and genera , Species Fungorum (in the Catalogue of Life) accepts 2 families, 56 genera, and 910 species in the Caliciales. *Caliciaceae ::''Acolium'' – 5 spp. ::'' Acroscyphus'' – 1 sp. ::''Allocalicium'' – 1 sp. ::''Amandinea'' – 83 spp. ::''Australiaena'' – 1 sp. ::''Baculifera'' – 18 spp. ::''Buellia'' – 201 spp. ::'' Caliciella'' – 1 sp. ::''Calicium'' – 36 spp. ::''Chrismofulvea'' – 3 spp. ::''Ciposia'' – 1 sp. ::''Cratiria'' – 23 spp. ::''Dermatiscum'' – 2 sp. ::''Dermiscellum'' – 1 sp. ::''Dimelaena'' – 10 spp. ::''Diploicia'' – 6 spp. ::''Diplotomma'' – 12 spp. ::''Dirinaria'' – 18 spp. ::''Endohyalina'' – 10 sp. ::'' Fluctua'' – 1 sp. ::''Gas ...
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Caliciales Genera
Caliciales is an order of mostly lichenized fungi in the class Lecanoromycetes. It consists of two families: Caliciaceae and Physciaceae, which together contain 54 genera and more than 1200 species. The order was circumscribed by American botanist Charles Edwin Bessey in 1907. Families and genera , Species Fungorum (in the Catalogue of Life) accepts 2 families, 56 genera, and 910 species in the Caliciales. *Caliciaceae ::''Acolium'' – 5 spp. ::'' Acroscyphus'' – 1 sp. ::''Allocalicium'' – 1 sp. ::''Amandinea'' – 83 spp. ::''Australiaena'' – 1 sp. ::''Baculifera'' – 18 spp. ::''Buellia'' – 201 spp. ::'' Caliciella'' – 1 sp. ::''Calicium'' – 36 spp. ::'' Chrismofulvea'' – 3 spp. ::'' Ciposia'' – 1 sp. ::'' Cratiria'' – 23 spp. ::'' Dermatiscum'' – 2 sp. ::'' Dermiscellum'' – 1 sp. ::'' Dimelaena'' – 10 spp. ::'' Diploicia'' – 6 spp. ::'' Diplotomma'' – 12 spp. ::'' Dirinaria'' – 18 spp. ::'' Endohyalina'' – 10 sp. ::'' Fluctua'' –  ...
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Wells Gray Provincial Park
Wells Gray Provincial Park is a large wilderness park located in east-central British Columbia, Canada. The park protects most of the southern, and highest, regions of the Cariboo Mountains and covers 5,250 square kilometres (524,990 hectares or 1.3 million acres). It is British Columbia's fourth largest park, after Tatshenshini-Alsek Provincial Park, Tatshenshini, Spatsizi Plateau Wilderness Provincial Park, Spatsizi and Tweedsmuir South Provincial Park, Tweedsmuir.Neave, Roland (2015). ''Exploring Wells Gray Park'', 6th edition. Wells Gray Tours, Kamloops, BC. .Goward, Trevor and Hickson, Cathie (1995). ''Nature Wells Gray'', 2nd edition. Lone Pine Publishing, Edmonton, AB. . Topography The boundaries of Wells Gray Park encompass 60 percent of the drainage basin of the Clearwater River (British Columbia), Clearwater River and most water that originates in the park flows into this river. The northern two-thirds of the park is extremely rugged with relief ranging from Clearwater L ...
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British Columbia
British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, forests, lakes, mountains, inland deserts and grassy plains, and borders the province of Alberta to the east and the Yukon and Northwest Territories to the north. With an estimated population of 5.3million as of 2022, it is Canada's third-most populous province. The capital of British Columbia is Victoria and its largest city is Vancouver. Vancouver is the third-largest metropolitan area in Canada; the 2021 census recorded 2.6million people in Metro Vancouver. The first known human inhabitants of the area settled in British Columbia at least 10,000 years ago. Such groups include the Coast Salish, Tsilhqotʼin, and Haida peoples, among many others. One of the earliest British settlements in the area was Fort Victoria, established ...
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Johannes Musaeus Norman
Johannes is a Medieval Latin form of the personal name that usually appears as "John" in English language contexts. It is a variant of the Greek and Classical Latin variants (Ιωάννης, ''Ioannes''), itself derived from the Hebrew name '' Yehochanan'', meaning "Yahweh is gracious". The name became popular in Northern Europe, especially in Germany because of Christianity. Common German variants for Johannes are ''Johann'', ''Hannes'', '' Hans'' (diminutized to ''Hänschen'' or ''Hänsel'', as known from "''Hansel and Gretel''", a fairy tale by the Grimm brothers), '' Jens'' (from Danish) and ''Jan'' (from Dutch, and found in many countries). In the Netherlands, Johannes was without interruption the most common masculine birth name until 1989. The English equivalent for Johannes is John. In other languages *Joan, Jan, Gjon, Gjin and Gjovalin in Albanian *'' Yoe'' or '' Yohe'', uncommon American form''Dictionary of American Family Names'', Oxford University Press, 2013. *Yaḥy ...
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Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus comes above species and below family (taxonomy), family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants ...
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Lichen
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

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Fungi
A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from the other eukaryotic kingdoms, which by one traditional classification include Plantae, Animalia, Protozoa, and Chromista. A characteristic that places fungi in a different kingdom from plants, bacteria, and some protists is chitin in their cell walls. Fungi, like animals, are heterotrophs; they acquire their food by absorbing dissolved molecules, typically by secreting digestive enzymes into their environment. Fungi do not photosynthesize. Growth is their means of mobility, except for spores (a few of which are flagellated), which may travel through the air or water. Fungi are the principal decomposers in ecological systems. These and other differences place fungi in a single group of related organisms, named the ''Eumycota'' (''t ...
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Monotypic
In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispecific" or "monospecific" is sometimes preferred. In botanical nomenclature, a monotypic genus is a genus in the special case where a genus and a single species are simultaneously described. In contrast, an oligotypic taxon contains more than one but only a very few subordinate taxa. Examples Just as the term ''monotypic'' is used to describe a taxon including only one subdivision, the contained taxon can also be referred to as monotypic within the higher-level taxon, e.g. a genus monotypic within a family. Some examples of monotypic groups are: Plants * In the order Amborellales, there is only one family, Amborellaceae and there is only one genus, '' Amborella'', and in this genus there is only one species, namely ''Amborella trichopoda. ...
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Circumscription (taxonomy)
In biological taxonomy, circumscription is the content of a taxon, that is, the delimitation of which subordinate taxa are parts of that taxon. If we determine that species X, Y, and Z belong in Genus A, and species T, U, V, and W belong in Genus B, those are our circumscriptions of those two genera. Another systematist might determine that T, U, V, W, X, Y, and Z all belong in genus A. Agreement on circumscriptions is not governed by the Codes of Zoological or Botanical Nomenclature, and must be reached by scientific consensus. A goal of biological taxonomy is to achieve a stable circumscription for every taxon. This goal conflicts, at times, with the goal of achieving a natural classification that reflects the evolutionary history of divergence of groups of organisms. Balancing these two goals is a work in progress, and the circumscriptions of many taxa that had been regarded as stable for decades are in upheaval in the light of rapid developments in molecular phylogenetics ...
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Lichen Genera
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (