Theosodon
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Theosodon
''Theosodon'' is an extinct genus of litoptern mammal from the Early to Middle Miocene of South America. Description ''Theosodon'' was long-legged with a long neck resembling modern llamas or guanacos. It was large for a litoptern, reaching up to in length and weighing up to . It had a long neck and tapir-like, three-toed feet, and like other litopterns and modern horses, tapirs and rhinos, it bore its weight on its middle toes. Extraordinarily, rather than having nostrils at the front of its head, ''Theosodon'' had its nostrils on the top of its snout, halfway between the forehead and the tip of the snout, and its nostrils pointed upwards rather than forwards, possibly as an adaptation for browsing on prickly vegetation. History and species ''Theosodon'' has been known since the 19th century, and by 1910 seven species had been described within the genus, all from the early Miocene Santa Cruz Formation in Argentina. The name ''Theosodon'' means "god tooth". The first fo ...
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Theosodon NT
''Theosodon'' is an extinct genus of litoptern mammal from the Early to Middle Miocene of South America. Description ''Theosodon'' was long-legged with a long neck resembling modern llamas or guanacos. It was large for a litoptern, reaching up to in length and weighing up to . It had a long neck and tapir-like, three-toed feet, and like other litopterns and modern horses, tapirs and rhinos, it bore its weight on its middle toes. Extraordinarily, rather than having nostrils at the front of its head, ''Theosodon'' had its nostrils on the top of its snout, halfway between the forehead and the tip of the snout, and its nostrils pointed upwards rather than forwards, possibly as an adaptation for browsing on prickly vegetation. History and species ''Theosodon'' has been known since the 19th century, and by 1910 seven species had been described within the genus, all from the early Miocene Santa Cruz Formation in Argentina. The name ''Theosodon'' means "god tooth". The first fo ...
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Laventan
The Laventan ( es, Laventense) age is a period of geologic time (13.8 to 11.8 Ma) within the Middle Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Colloncuran and precedes the Mayoan age.Madden et al., 1997 Etymology The age is named after the Miocene Lagerstätte La Venta, where a rich biodiversity from the Middle Miocene has been recovered from the Honda Group. Formations Fossil content Correlations The Laventan (13.8 to 11.8 Ma) correlates with: * NALMA ** latest Barstovian (15.97-13.65 Ma)Barstovian
at Fossilworks.org
** early



Pternoconius
''Pternoconius'' is an extinct genus of macraucheniid litoptern from the Late Oligocene and Early Miocene of Argentina. Fossils of this genus have been found in the Sarmiento Formation of Argentina. Etymology The genus name, ''Pternoconius'', is an anagram of the closely related genus '' Coniopternium''. The species name refers to the similarity of the Eocene genus '' Polymorphis''. Species ''Pternoconius tournoueri'' In 1985, fossils found in the Early Miocene Colhué Huapí Member of the Sarmiento Formation were assigned to the genus under the name ''Pternoconius tournoueri'', consisting of a nearly complete hemimandible. ''Pternoconius bondi'' In 2016 a new species of macraucheniid litoptern was described coming from the Bajada del Diablo locality in the Sarmiento Formation, consisting of the anterior portion of the skull with the maxillary region, some fragments of nasal bones, a small portion of the left zygomatic process, and most of the upper dentition (i.e., l ...
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Cramauchenia
''Cramauchenia'' is an extinct genus of litoptern South American ungulate. ''Cramauchenia'' was named by Florentino Ameghino. The name has no literal translation. Instead, it is an anagram of the name of a related genus ''Macrauchenia''. This genus was initially discovered in the Sarmiento Formation in the Chubut Province, in Argentina, and later it was found in the Chichinales Formation in the Río Negro Province and the Cerro Bandera Formation in Neuquén, also in Argentina, in sediments assigned to the SALMA Colhuehuapian (in the Early Miocene), as well as the Agua de la Piedra Formation in Mendoza, in sediments dated to the Deseadan (during the Late Oligocene). In 1981 Soria made ''C. insolita'' a junior synonym of ''C. normalis''. A specimen of ''C. normalis'' was described in 2010 from Cabeza Blanca (Chubut, Argentina) in the Sarmiento Formation, in sediments assigned to the Deseadan SALMA (Upper Oligocene). Description This animal had an appearance vaguely similar ...
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Xenorhinotherium
''Xenorhinotherium'' is an extinct genus of macraucheniine macraucheniids, closely related to '' Macrauchenia'' of Patagonia. The type species is ''X. bahiense''.''Xenorhinotherium''
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Taxonomy

Some authors consider the genus ''Xenorhinotherium'' a synonym of ''Macrauchenia'', while all others consider it a distinct genus. The name ''Xenorhinotherium'' means "Strange-Nosed Beast" and ''bahiense'' refers to the Brazilian state of , where the first fossils were found. ''Xenorhinotherium'' wa ...
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Promacrauchenia
''Promacrauchenia'' is an extinct genus of macraucheniids that lived during the Late Miocene to Late Pliocene epochs of what is now Argentina and Bolivia. It belongs to the subfamily Macraucheniinae, which also includes ''Huayqueriana'', ''Macrauchenia'', and ''Xenorhinotherium.'' Fossils of this genus have been found in the Ituzaingó, Andalhuala, and Cerro Azul Formation The Cerro Azul Formation ( es, Formación Cerro Azul), in the Buenos Aires Province also described as Epecuén Formation, is a geological formation of Late Miocene (Tortonian, or Huayquerian in the SALMA classification) age in the Colorado Basin ...s of Argentina. Classification The genus ''Promacrauchenia'' was first described by Florentino Ameghino in 1904, on the basis of fossils found in Patagonia in lower Pliocene deposits and which Ameghino himself, years earlier, had described as a species of ''Macrauchenia'', as ''M. antiqua'' . In addition to the type species, ''Promacrauchenia'' ''antiqua'', o ...
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Cullinia
''Cullinia'' is an extinct genus of litoptern, an order of South American native ungulates that included horse-like and camel-like animals such as ''Macrauchenia''. It is only known from fragmentary remains. ''Cullinia levis'' is known from Chasicoan remains found in the Arroyo Chasicó Formation of Argentina, and remains from the Brazilian state of Acre and the Huayquerian Ituzaingó Formation have been assigned to ''Cullinia'' sp. . History and naming ''Cullinia'' was described in 1931 by Cabrera and Kraglievich, from MLP 29-IX-178, an holotype containing several fragmentary remains including a mandible and a metatarsal. In 1995, Bond and López add to the holotype other remains from the upper dentition. It was named from the Araucanian word "cullin", meaning "animal". Description ''Cullinia'' was a slender, small Macraucheniidae. It had proportionally larger metapodials than '' Theosodon'', and its first lower molar was absent. Classification Below is a phylogeneti ...
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Macraucheniidae
Macraucheniidae is a family in the extinct South American ungulate order Litopterna, that resembled various camelids. The reduced nasal bones of their skulls was originally suggested to have housed a small proboscis, similar to that of the saiga antelope. However, one study suggested that they were openings for large moose-like nostrils. Conversely, prehistoric pictographs by indigenous people seems to depict animals interpreted as macraucheniids with trunks. Their hooves were similar to those of rhinoceroses today, with a simple ankle joint and three digits on each foot. Thus, they may have been capable of rapid directional change when running away from predators, such as large phorusrhacid terror birds, sparassodont metatherians, giant short-faced bears ('' Arctotherium)'' and saber-toothed cats (''Smilodon''). Macraucheniids probably lived in large herds to gain protection against these predators, as well as to facilitate finding mates for reproduction. The family Macrauc ...
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Scalabrinitherium
''Scalabrinitherium'' is an extinct genus of mammals of the family Macraucheniidae. Fossils of this animal were found among the fossils of prehistoric xenarthrans in the Ituzaingó Formation of Argentina.''Scalabrinitherium''
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Description

This animal was rather similar to a with a slightly heavy build; the was long and low, the front teeth were slightly spatulate, and the nasal aperture set far back. It is possible that there was a strong prehens ...
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Santa Cruz Formation, Argentina
The Santa Cruz Formation is a geological formation in the Magallanes/Austral Basin in southern Patagonia in Argentina and in adjacent areas of Chile. It dates to the late Early Miocene epoch, and is contemporaneous with eponymous Santacrucian SALMA. The formation extends from the Andes to the Atlantic coast. In its coastal section it is divided into two members, the lower, fossil rich Estancia La Costa Member, which has a lithology predominantly consisting of tuffaceous deposits and fine grained sedimentary Mudrock, claystone and mudstone, and the upper fossil-poor Estancia La Angelina Member, which consists of sedimentary rock, primarily claystone, mudstone, and sandstone. The environment of deposition is interpreted to have been mostly Fluvial processes, fluvial, with the lowermost part of the Estancia La Costa Member being transitional between fluvial and marine conditions. The environment of the Estancia La Costa Member is thought to have been relatively warm and humid, but ...
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Windhausenia
''Windhausenia'' is an extinct genus of mammals belonging to the family Macraucheniidae and the order Litopterna. While it reached the size of its better known relative ''Macrauchenia'', its constitution was lighter. Remains from the genus have been uncovered in Argentina. ''Windhausenia'' fossils were only found in the middle layers of the Uquía Formation. The genus was described in 1930 by Kraglievich, who considered it more derived than '' Promacrauchenia'', but less than ''Macrauchenia''. The genus survived the Great American Interchange. It had to compete with more derived relatives such as ''Macrauchenia'', and may have occupied a specialised ecological niche to avoid competition. Its remains are found in subtropical areas, while other genera were found in tropical and temperate environment. As fossils of this genus are often found in association with aeolian deposits, corresponding with arid and semiarid environments, it is possible it occupied more similar with m ...
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Huayqueriana
''Huayqueriana'' is an extinct genus of South American litoptern, related to ''Macrauchenia'', and belonging to the same family, Macraucheniidae. It was formerly known as ''Macrauchenidia latidens'', described in 1939 by Cabrera, but redefined as ''Huayqueriana'' in 2016 based on the earlier name convention of Rovereto 1914.Forasiepi et al., 2016, p.11Cabrera (A.), 1939. Sobre vertebrados fósiles del Plioceno del Adolfo Alsina. ''Revista Museo de La Plata'', II (n. Serie), Paleont.: 3-35. The genus is named after the Huayquerías Formation and the eponymous Huayquerian South American land mammal age defined at the formation. Classification Cladogram based in the phylogenetic analysis In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ... published by Schmidt ''et al''., 2014, showi ...
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