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Pternoconius
''Pternoconius'' is an extinct genus of macraucheniid litoptern from the Late Oligocene and Early Miocene of Argentina. Fossils of this genus have been found in the Sarmiento Formation of Argentina. Etymology The genus name, ''Pternoconius'', is an anagram of the closely related genus '' Coniopternium''. The species name refers to the similarity of the Eocene genus '' Polymorphis''. Species ''Pternoconius tournoueri'' In 1985, fossils found in the Early Miocene Colhué Huapí Member of the Sarmiento Formation were assigned to the genus under the name ''Pternoconius tournoueri'', consisting of a nearly complete hemimandible. ''Pternoconius bondi'' In 2016 a new species of macraucheniid litoptern was described coming from the Bajada del Diablo locality in the Sarmiento Formation, consisting of the anterior portion of the skull with the maxillary region, some fragments of nasal bones, a small portion of the left zygomatic process, and most of the upper dentition (i.e., l ...
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Cramauchenia
''Cramauchenia'' is an extinct genus of litoptern South American ungulate. ''Cramauchenia'' was named by Florentino Ameghino. The name has no literal translation. Instead, it is an anagram of the name of a related genus ''Macrauchenia''. This genus was initially discovered in the Sarmiento Formation in the Chubut Province, in Argentina, and later it was found in the Chichinales Formation in the Río Negro Province and the Cerro Bandera Formation in Neuquén, also in Argentina, in sediments assigned to the SALMA Colhuehuapian (in the Early Miocene), as well as the Agua de la Piedra Formation in Mendoza, in sediments dated to the Deseadan (during the Late Oligocene). In 1981 Soria made ''C. insolita'' a junior synonym of ''C. normalis''. A specimen of ''C. normalis'' was described in 2010 from Cabeza Blanca (Chubut, Argentina) in the Sarmiento Formation, in sediments assigned to the Deseadan SALMA (Upper Oligocene). Description This animal had an appearance vaguely similar ...
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Sarmiento Formation
The Sarmiento Formation (Spanish: ''Formación Sarmiento''), in older literature described as the Casamayor Formation, is a geological formation in Chubut Province, Argentina, in central Patagonia, which spans around 30 million years from the mid-Eocene to the early Miocene. It predominantly consists of pyroclastic deposits, which were deposited in a semi-arid environment. It is divided up into a number of members. The diverse fauna of the Sarmiento Formation, including a variety of birds, crocodilians, turtles and snakes, also includes many mammals such as South American native ungulates (notoungulates, litopterns, astrapotheres) as well as armadillo Armadillos (meaning "little armored ones" in Spanish) are New World placental mammals in the order Cingulata. The Chlamyphoridae and Dasypodidae are the only surviving families in the order, which is part of the superorder Xenarthra, along wi ...s, and caviomorph rodents. Material was copied from this source, which is availabl ...
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Colhuehuapian
The Colhuehuapian age is a period of geologic time (21.0–17.5 Ma) within the Early Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Deseadan and precedes the Santacrucian age. Etymology The age is named after the Colhué Huapí Member of the Sarmiento Formation in the Golfo San Jorge Basin, Patagonia, Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th .... Formations Fossils References Bibliography ;Colhué Huapí Member * * ;Abanico Formation * * * * * * ;Biblián Formation * ;Castillo Formation * * * * * ;Cerro Bandera Formation * * ;Chichinales Formation * * ;Chilcatay Formation * * * * * * * * * * ;Cura-Mallín Group * * * ;Gaiman Formation * ...
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Cramaucheniinae
Cramaucheniinae is a paraphyletic subfamily of macraucheniids that originated in the middle Eocene (Mustersan SALMA). The size range of the group ranged from small, basal forms to larger and more derived forms. During their evolution, the cramaucheniines undergone a trend from evolving from small basal forms such as '' Polymorphis'' into larger, more derived taxa such as ''Theosodon ''Theosodon'' is an extinct genus of litoptern mammal from the Early to Middle Miocene of South America. Description ''Theosodon'' was long-legged with a long neck resembling modern llamas or guanacos. It was large for a litoptern, reaching ...''. References Macraucheniids Paleogene mammals of South America Neogene mammals of South America Mammal subfamilies Lutetian first appearances Serravallian extinctions Paraphyletic groups {{Paleo-mammal-stub ...
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Paranauchenia
''Paranauchenia'' is an extinct genus of South American litopterns belonging to the family Macraucheniidae. It is known only from fossil finds in Argentina. It possessed three toes and long limbs. The species ''Paranauchenia denticulata'' lived in the Miocene epoch in Argentina.Ameghino, F. 1904a. Nuevas especies de mamíferos Cretáceos y Terciarios de la República Argentina. ''Anales de la Sociedad Científica Argentina'' 16: 193-208; 17: 162-175, 182-192, 225-240, 241-291. Fossils have been found in the Arroyo Chasicó and Ituzaingó Formations of Argentina.''Paranauchenia''
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Theosodon
''Theosodon'' is an extinct genus of litoptern mammal from the Early to Middle Miocene of South America. Description ''Theosodon'' was long-legged with a long neck resembling modern llamas or guanacos. It was large for a litoptern, reaching up to in length and weighing up to . It had a long neck and tapir-like, three-toed feet, and like other litopterns and modern horses, tapirs and rhinos, it bore its weight on its middle toes. Extraordinarily, rather than having nostrils at the front of its head, ''Theosodon'' had its nostrils on the top of its snout, halfway between the forehead and the tip of the snout, and its nostrils pointed upwards rather than forwards, possibly as an adaptation for browsing on prickly vegetation. History and species ''Theosodon'' has been known since the 19th century, and by 1910 seven species had been described within the genus, all from the early Miocene Santa Cruz Formation in Argentina. The name ''Theosodon'' means "god tooth". The first fo ...
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Scalabrinitherium
''Scalabrinitherium'' is an extinct genus of mammals of the family Macraucheniidae. Fossils of this animal were found among the fossils of prehistoric xenarthrans in the Ituzaingó Formation of Argentina.''Scalabrinitherium''
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Description

This animal was rather similar to a with a slightly heavy build; the was long and low, the front teeth were slightly spatulate, and the nasal aperture set far back. It is possible that there was a strong prehens ...
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Huayqueriana
''Huayqueriana'' is an extinct genus of South American litoptern, related to ''Macrauchenia'', and belonging to the same family, Macraucheniidae. It was formerly known as ''Macrauchenidia latidens'', described in 1939 by Cabrera, but redefined as ''Huayqueriana'' in 2016 based on the earlier name convention of Rovereto 1914.Forasiepi et al., 2016, p.11Cabrera (A.), 1939. Sobre vertebrados fósiles del Plioceno del Adolfo Alsina. ''Revista Museo de La Plata'', II (n. Serie), Paleont.: 3-35. The genus is named after the Huayquerías Formation and the eponymous Huayquerian South American land mammal age defined at the formation. Classification Cladogram based in the phylogenetic analysis In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ... published by Schmidt ''et al''., 2014, showi ...
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Litoptern
Litopterna (from grc, λῑτή πτέρνα "smooth heel") is an extinct order of fossil hoofed mammals from the Cenozoic era. The order is one of the five great orders of South American ungulates that were endemic to the continent, until the Great American Biotic Interchange brought new ungulate species. Like other endemic South American mammals, their relationship to other mammal groups had long been unclear, but recent genetic and proteomic evidence indicates that their closest living relatives are Perissodactyls (odd-toed ungulates) including horses, rhinoceros, and tapirs, and that litopterns are closely related to notoungulates, another widespread group of South American ungulates. There were two major groups of litopterns: Proterotheriidae and Macraucheniidae. Proterotheriids were medium to large animals that evolved adaptations for fast running, and occupied a variety of niches that elsewhere were filled by animals such as goats and antelopes, mouse deer, and hors ...
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Cullinia
''Cullinia'' is an extinct genus of litoptern, an order of South American native ungulates that included horse-like and camel-like animals such as ''Macrauchenia''. It is only known from fragmentary remains. ''Cullinia levis'' is known from Chasicoan remains found in the Arroyo Chasicó Formation of Argentina, and remains from the Brazilian state of Acre and the Huayquerian Ituzaingó Formation have been assigned to ''Cullinia'' sp. . History and naming ''Cullinia'' was described in 1931 by Cabrera and Kraglievich, from MLP 29-IX-178, an holotype containing several fragmentary remains including a mandible and a metatarsal. In 1995, Bond and López add to the holotype other remains from the upper dentition. It was named from the Araucanian word "cullin", meaning "animal". Description ''Cullinia'' was a slender, small Macraucheniidae. It had proportionally larger metapodials than '' Theosodon'', and its first lower molar was absent. Classification Below is a phylogeneti ...
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Promacrauchenia
''Promacrauchenia'' is an extinct genus of macraucheniids that lived during the Late Miocene to Late Pliocene epochs of what is now Argentina and Bolivia. It belongs to the subfamily Macraucheniinae, which also includes ''Huayqueriana'', ''Macrauchenia'', and ''Xenorhinotherium.'' Fossils of this genus have been found in the Ituzaingó, Andalhuala, and Cerro Azul Formation The Cerro Azul Formation ( es, Formación Cerro Azul), in the Buenos Aires Province also described as Epecuén Formation, is a geological formation of Late Miocene (Tortonian, or Huayquerian in the SALMA classification) age in the Colorado Basin ...s of Argentina. Classification The genus ''Promacrauchenia'' was first described by Florentino Ameghino in 1904, on the basis of fossils found in Patagonia in lower Pliocene deposits and which Ameghino himself, years earlier, had described as a species of ''Macrauchenia'', as ''M. antiqua'' . In addition to the type species, ''Promacrauchenia'' ''antiqua'', o ...
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Windhausenia
''Windhausenia'' is an extinct genus of mammals belonging to the family Macraucheniidae and the order Litopterna. While it reached the size of its better known relative ''Macrauchenia'', its constitution was lighter. Remains from the genus have been uncovered in Argentina. ''Windhausenia'' fossils were only found in the middle layers of the Uquía Formation. The genus was described in 1930 by Kraglievich, who considered it more derived than '' Promacrauchenia'', but less than ''Macrauchenia''. The genus survived the Great American Interchange. It had to compete with more derived relatives such as ''Macrauchenia'', and may have occupied a specialised ecological niche to avoid competition. Its remains are found in subtropical areas, while other genera were found in tropical and temperate environment. As fossils of this genus are often found in association with aeolian deposits, corresponding with arid and semiarid environments, it is possible it occupied more similar with m ...
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