Thalassocnus
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Thalassocnus
''Thalassocnus'' is an extinct genus of semiaquatic ground sloths from the Miocene and Pliocene of the Pacific South American coast. It is monotypic within the subfamily Thalassocninae. The five species—''T. antiquus'', ''T. natans'', ''T. littoralis'', ''T. carolomartini'', and ''T. yuacensis''—represent a chronospecies, a population gradually adapting to marine life in one direct lineage. They are the only known aquatic sloths, but they may have also been adapted to a terrestrial lifestyle. They have been found in the Pisco Formation of Peru, the Tafna Formation of Argentina, and the Bahía Inglesa, Coquimbo, and Horcón formations of Chile. Thalassocninae has been placed in both the families Megatheriidae and Nothrotheriidae. ''Thalassocnus'' evolved several marine adaptations over the course of 4 million years, such as dense and heavy bones to counteract buoyancy, the internal nostrils migrating farther into the head to help with breathing while completely submerged, ...
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Pisco Formation
The Pisco Formation is a geologic formation located in Peru, on the southern coastal desert of Ica, Peru, Ica and Arequipa. The approximately thick formation was deposited in the Pisco Basin, spanning an age from the Miocene, Middle Miocene up to the Early Pleistocene, roughly from 15 to 2 Ma. The tuffaceous sandstones, diatomaceous siltstones, Conglomerate (geology), conglomerates and dolomite (rock), dolomites were deposited in a lagoonal to near-shore depositional environment, environment, in bays similar to other Pacific South American formations as the Caldera Basin, Bahía Inglesa and Coquimbo Formations of Chile. Several specialists consider the Pisco Formation one of the most important Lagerstätten,Brand et al., 2004Brand et al., 2011 based on the large amount of exceptionally preserved marine fossils, including sharks (most notably megalodon), penguins, whales, dolphins, birds, marine crocodiles and aquatic giant sloths. Famous fossils found in these layers include the ...
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Nothrotheriidae
Nothrotheriidae is a family of extinct ground sloths that lived from approximately 17.5 mya—10,000 years ago, existing for approximately . Previously placed within the tribe Nothrotheriini or subfamily Nothrotheriinae within Megatheriidae, they are now usually placed in their own family, Nothrotheriidae. Nothrotheriids appeared in the Burdigalian, some 19.8 million years ago, in South America. The group includes the comparatively slightly built ''Nothrotheriops'', which reached a length of about . While nothrotheriids were small compared to some of their megatheriid relatives, their claws provided an effective defense against predators, like those of larger anteaters today. Evolution During the late Miocene and Pliocene, the sloth genus ''Thalassocnus'' of the west coast of South America became adapted to a shallow-water marine lifestyle. However, the family placement of ''Thalassocnus'' has been disputed; while long considered a nothrotheriid, one recent analysis moves it to ...
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Megatheriidae
Megatheriidae is a family of extinct ground sloths that lived from approximately 23 mya—11,000 years ago. Megatheriids appeared during the Late Oligocene (Deseadan in the SALMA classification), some 29 million years ago, in South America. The group includes the heavily built '' Megatherium'' (given its name 'great beast' by Georges Cuvier) and ''Eremotherium''. An early genus that was originally considered a megatheriid, the more slightly built ''Hapalops'', reached a length of about . The nothrotheres have recently been placed in their own family, Nothrotheriidae. The skeletal structure of these ground sloths indicates that the animals were massive. Their thick bones and even thicker joints (especially those on the hind legs) gave their appendages tremendous power that, combined with their size and fearsome claws, provided a formidable defense against predators. The earliest megatheriid in North America was ''Sibotherium'' which arrived 5.3 million years ago, after cros ...
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South American Land Mammal Age
The South American land mammal ages (SALMA) establish a geologic timescale for prehistoric South American fauna beginning 64.5 Mya (unit), Ma during the Paleocene and continuing through to the Late Pleistocene (0.011 Ma). These periods are referred to as ages, stages, or intervals and were established using geographic place names where fossil materials where obtained.Flynn & Swisher, 1995 The basic unit of measure is the first/last boundary statement. This shows that the first appearance event of one taxon is known to predate the last appearance event of another. If two taxa are found in the same fossil quarry or at the same stratigraphic horizon, then their age-range zones overlap. Background South America was an island continent for much of the Cenozoic, or the "Age of Mammals". As a result, its mammals evolved in their own unique directions, as Australia and Madagascar still have today. Paleogeographic timeline A simplified paleogeography, paleogeographic timeline of South ...
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Huayquerian
The Huayquerian ( es, Huayqueriense) age is a period of geologic time (9.0–6.8 Ma) within the Late Miocene epoch of the Neogene, used more specifically within the SALMA classification. It follows the Mayoan and precedes the Montehermosan age. Etymology The age is named after the Huayquerías Formation in the western Cuyo Basin of northwestern Argentina that was later dated to the Montehermosan. The most complete Huayquerian fauna is found in the Cerro Azul Formation The Cerro Azul Formation ( es, Formación Cerro Azul), in the Buenos Aires Province also described as Epecuén Formation, is a geological formation of Late Miocene (Tortonian, or Huayquerian in the SALMA classification) age in the Colorado Basin ..., in Buenos Aires Province also referred to as Epecuén Formation. Formations Fossil content Correlations Notes and references Notes References Bibliography ;Huayquerías Formation * * ;Andalhuala Formation * ;Camacho Formation * * ...
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Acrophyseter
''Acrophyseter'' is a genus of extinct sperm whales that lived in the Late Miocene off the coast of Peru comprising two species: ''A. deinodon'' and ''A. robustus''. It is part of a group of macroraptorial sperm whales which all shared several features for the purpose of hunting large prey, such as deeply-rooted and thick teeth. ''Acrophyseter'' measured , making it the smallest raptorial sperm whale. Because of its short pointed snout, and its strong curved front teeth, it probably fed on the large marine vertebrates of its time, such as seals and other whales. Discovery The genus ''Acrophyseter'' houses two species. The type species, ''A. deinodon'', was discovered in the Sud-Sacaco locality of the Pisco Formation in Peru, dating back to the Tortonian–Messinian stages of the Miocene around 8.5–6.7 million years ago (mya); the holotype specimen, MNHN SAS 1626, represents a mature individual and consists of a skull and jaw with most of the teeth intact. The second species, '' ...
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Ground Sloth
Ground sloths are a diverse group of extinct sloths in the mammalian superorder Xenarthra. The term is used to refer to all extinct sloths because of the large size of the earliest forms discovered, compared to existing tree sloths. The Caribbean ground sloths, the most recent survivors, lived in the Antilles, possibly until 1550 BCE. However, radiocarbon dating suggests an age of between 2819 and 2660 BCE for the last occurrence of ''Megalocnus'' in Cuba. Ground sloths had been extinct on the mainland of North and South America for 10,000 years or more. They survived 5,000–6,000 years longer in the Caribbean than on the American mainland, which correlates with the later colonization of this area by humans. Much ground sloth evolution took place during the late Paleogene and Neogene of South America, while the continent was isolated. At their earliest appearance in the fossil record, the ground sloths were already distinct at the family level. The presence ...
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Chronospecies
A chronospecies is a species derived from a anagenesis, sequential development pattern that involves continual and uniform changes from an extinct ancestral form on an evolutionary scale. The sequence of alterations eventually produces a population that is physically, Comparative anatomy, morphologically, and/or Genetics, genetically distinct from the original ancestors. Throughout the change, there is only one species in the lineage at any point in time, as opposed to cases where divergent evolution produces contemporary species with a common ancestor. The related term paleospecies (or palaeospecies) indicates an extinct species only identified with fossil material. That identification relies on distinct similarities between the earlier fossil specimens and some proposed descendant although the exact relationship to the later species is not always defined. In particular, the range of variation within all the early fossil specimens does not exceed the observed range that exists in t ...
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Marine Mammal
Marine mammals are aquatic mammals that rely on the ocean and other marine ecosystems for their existence. They include animals such as seals, whales, manatees, sea otters and polar bears. They are an informal group, unified only by their reliance on marine environments for feeding and survival. Marine mammal adaptation to an aquatic lifestyle varies considerably between species. Both cetaceans and sirenians are fully aquatic and therefore are obligate water dwellers. Seals and sea-lions are semiaquatic; they spend the majority of their time in the water but need to return to land for important activities such as mating, breeding and molting. In contrast, both otters and the polar bear are much less adapted to aquatic living. The diets of marine mammals vary considerably as well; some eat zooplankton, others eat fish, squid, shellfish, or sea-grass, and a few eat other mammals. While the number of marine mammals is small compared to those found on land, their roles in various ...
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Macroraptorial Sperm Whale
Macroraptorial sperm whales were highly predatory whales of the sperm whale superfamily (Physeteroidea) of the Miocene epoch that hunted large marine mammals, including other whales, using their large teeth. They consist of five genera: ''Acrophyseter'', ''Albicetus'', ''Brygmophyseter'', ''Livyatan'', and ''Zygophyseter''. All species are known by at least a skull, and are informally grouped without a family designation. They were all likely the apex predator of their habitats, comparable to the modern day killer whale (''Orcinus orca''), and achieved great lengths, with one species–''Livyatan''–measuring about . Discovery ''Zygophyseter'' was discovered in the Pietra Leccese Formation in Italy from a skull, teeth, and vertebrae; ''Brygmophyseter'' was discovered in the Bessho Formation in Japan from a nearly-complete skeleton; and ''Acrophyseter'' and ''Livyatan'' both originate from the Pisco Formation in Peru and are known by only a skull. ''Albicetus'' is discovered fro ...
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Seagrass
Seagrasses are the only flowering plants which grow in marine environments. There are about 60 species of fully marine seagrasses which belong to four families (Posidoniaceae, Zosteraceae, Hydrocharitaceae and Cymodoceaceae), all in the order Alismatales (in the clade of monocotyledons). Seagrasses evolved from terrestrial plants which recolonised the ocean 70 to 100 million years ago. The name ''seagrass'' stems from the many species with long and narrow leaves, which grow by rhizome extension and often spread across large "meadows" resembling grassland; many species superficially resemble terrestrial grasses of the family Poaceae. Like all autotrophic plants, seagrasses photosynthesize, in the submerged photic zone, and most occur in shallow and sheltered coastal waters anchored in sand or mud bottoms. Most species undergo submarine pollination and complete their life cycle underwater. While it was previously believed this pollination was carried out without pollinators ...
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Late Miocene
The Late Miocene (also known as Upper Miocene) is a sub-epoch of the Miocene epoch (geology), Epoch made up of two faunal stage, stages. The Tortonian and Messinian stages comprise the Late Miocene sub-epoch, which lasted from 11.63 Ma (million years ago) to 5.333 Ma. The evolution of life The gibbons (family Hylobatidae) and orangutans (genus ''Pongo'') are the first groups to split from the line leading to the hominins, including humans, then gorillas (genus ''Gorilla''), and finally, chimpanzees and bonobos (genus ''Pan (genus), Pan''). The splitting date between hominin and chimpanzee lineages is placed by some between 4 to 8 million years ago, that is, during the Late Miocene. References External links GeoWhen Database - Late Miocene
Miocene, .03 Miocene geochronology, 03 Messinian, * Tortonian, * {{geochronology-stub ...
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